Hallucigenia is a genus of Cambrian animal resembling worms, known from articulated fossils in Burgess Shale-type deposits in Canada and China, and from isolated spines around the world. The generic name reflects the type species' unusual appearance and eccentric history of study; when it was erected as a genus, H. sparsa was reconstructed as an enigmatic animal upside down and back to front. Hallucigenia was later recognized as a lobopodian, a grade of Paleozoic panarthropods from which the velvet worms, water bears, and arthropods arose.
Parapeytoia was a prehistoric arthropod that lived over 530 million years ago in the Maotianshan shales of prehistoric China. It was interpreted as an anomalocaridid (radiodont) with legs, but later studies reveal it was a megacheiran, a group of arthropods which are no longer thought to be closely related to the radiodonts.
Mixopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Mixopterus have been discovered in deposits from Late Silurian age, and have been referred to several different species. Fossils have been recovered from two continents; Europe and North America.
Acanthomeridion is an extinct arthropod found in the Chengjiang fauna deposits of China. In 1997, it was placed in its own, monotypic family, Acanthomeridiidae. It is known from eight specimens, all found in China.
Megacheira is an extinct class of predatory arthropods that possessed a pair of great appendages, hence the class name as well as the common name "great appendage arthropods". Their taxonomic position is controversial, with studies either considering them stem-group euarthropods, or stem-group chelicerates. Most of them were found in marine environments throughout the world from the lower to middle Cambrian. Megacheirans were important components of several faunas, including the Burgess, Wheeler and Maotianshan Shales Lagerstatten.
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xiphosura sensu stricto and only regarded as horseshoe crabs under a broader sense. Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata.
Cucumericrus decoratus is a species of putative radiodont known from a few poorly preserved specimens. Only fragments of trunk cuticle and corresponded appendages had been revealed, while important radiodont features such as frontal appendages are unknown in this species. The trunk cuticle possess irregular wrinkles and may had been soft in life. Each of the trunk appendage compose of a dorsal flap-like element and a ventral stubby leg with unknown distal region, structurally comparable to the trunk appendages of gill lobopodians and euarthropod biramous appendages. The legs have been interpreted as somewhere between annulated lobopod legs and segmented arthropod legs.
Pauline avibella is a fossil ostracod from the Silurian with unusually well preserved soft parts, including limbs, eyes, gills and alimentary system.
Pauline is a fossil genus of ostracods from the Silurian. Genus contains two species: Pauline avibella found in 425-million-year-old rocks in the Herefordshire Lagerstätte in England near the Welsh Border and Pauline nivisis, known from the Lower Silurian Pentamerus Bjerge Formation of north Greenland.
Aquilonifer spinosus is an extinct species of arthropod from the Silurian period. It is known from a single fossil specimen found in the Wenlock Series Lagerstätte of Herefordshire, England, in rocks about 430 million years old. The 1 cm long specimen is a stem-group mandibulate, not directly related to any living species. The many-legged, eyeless adult has ten unusual tethered appendages, interpreted as juveniles attached to the parent, in a unique form and previously unknown brooding behaviour.
Prosomapoda is a clade of euchelicerates including the groups Xiphosura and Planaterga, as well as several basal synziphosurid genera. The clade is defined by the lack of exopods of prosomal appendage II-V in the adult instar, where in contrast the exopods of appendage II-V are well-developed in the non-prosomapod euchelicerates Offacolus and Dibasterium.
Offacolus is an extinct genus of euchelicerate, a group of chelicerate arthropods. Its only species, O. kingi, has been found in deposits from the Silurian period in the Wenlock Series Lagerstätte of Herefordshire, England. It is the only member of the monotypic family Offacolidae, and classified as a basal ("primitive") genus in the clade Euchelicerata, along with Dibasterium and Prosomapoda. The genus is named after Offa, a king from the ancient kingdom of Mercia, and colus, a person who dwelled among the Offa's Dyke. The species name honors Robert Joseph King, a British mineralogist who found the fossils of Offacolus.
Palaeoisopus is a monotypic genus of fossil pycnogonid, known only by one species, Palaeoisopus problematicus, discovered from the Lower Devonian Hunsrück Slate of Germany. It have several characters unusual for a pycnogonid, such as swimming legs with alternating size, medially-arranged eyes, and most significantly, a long, segmented abdomen, which were highly reduced in modern counterparts.
Luolishania is an extinct genus of lobopodian worm and known from the Lower Cambrian Chiungchussu Formation of the Chengjiang County, Yunnan Province, China. A monotypic genus, it contains one species Luolishania longicruris. It was discovered and described by Hou Xian-Guang and Chen Jun-Yuan in 1989. It is one of the superarmoured Cambrian lobopodians suspected to be either an intermediate form in the origin of velvet worms (Onychophora) or basal to at least Tardigrada and Arthropoda. It is the basis of the family name Luolishaniidae, which also include other related lobopods such as Acinocricus, Collinsium, Facivermis, and Ovatiovermis. Along with Microdictyon, it is the first lobopodian fossil discovered from China.
Thanahita is a genus of extinct lobopodian and known from the middle Silurian Herefordshire Lagerstätte at the England–Wales border in UK. It is monotypic and contains one species, Thanahita distos. Discovered in 2018, it is estimated to have lived around 430 million years ago and is the only known extinct lobopodian in Europe, and the first Silurian lobopodian known worldwide.
Hallucigeniidae is a family of extinct worms belonging to the group Lobopodia that originated during the Cambrian explosion. It is based on the species Hallucigenia sparsa, the fossil of which was discovered by Charles Doolittle Walcott in 1911 from the Burgess Shale of British Columbia. The name Hallucigenia was created by Simon Conway Morris in 1977, from which the family was erected after discoveries of other hallucigeniid worms from other parts of the world. Classification of these lobopods and their retatives are still controversial, and the family consists of at least four genera.
Coalbrookdale Formation, earlier known as Wenlock Shale or Wenlock Shale Formation and also referred to as Herefordshire Lagerstätte in palaeontology, is a fossil-rich deposit (Konservat-Lagerstätte) in Powys and Herefordshire at the England–Wales border in UK. It belongs to the Wenlock Series of the Silurian Period within the Homerian Age. It is known for its well-preserved fossils of various invertebrate animals many of which are in their three-dimensional structures. Some of the fossils are regarded as earliest evidences and evolutionary origin of some of the major groups of modern animals.
Carbotubulus is a genus of extinct worm belonging to the group Lobopodia and known from the Carboniferous Carbondale Formation of the Mazon Creek area in Illinois, US. A monotypic genus, it contains one species Carbotubulus waloszeki. It was discovered and described by Joachim T. Haug, Georg Mayer, Carolin Haug, and Derek E.G. Briggs in 2012. With an age of about 300 million years, it is the first long-legged lobopodian discovered after the period of Cambrian explosion.
Enalikter is an extinct arthropod described from the middle Silurian Herefordshire Lagerstätte at the England–Wales border in UK. This genus is known from only one species, E. aphson. Enalikter is described as late-living example of Megacheira, "great-appendage arthropod". It was once considered as an annelid by other researchers, however subsequent studies denied this. Its interpretation as megacheiran arthropod is questioned in later study.
Bundenbachiellus is an extinct genus of arthropod described from the Lower Devonian Hunsrück Slate of Germany. This genus is known from only one species, B. giganteus. Alongside its possible relative Enalikter from Silurian, it is possible that genus is late-living example of Megacheira, "great-appendage arthropod".
