Dociostaurus maroccanus | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Orthoptera |
Suborder: | Caelifera |
Family: | Acrididae |
Subfamily: | Gomphocerinae |
Tribe: | Dociostaurini |
Genus: | Dociostaurus |
Species: | D. maroccanus |
Binomial name | |
Dociostaurus maroccanus (Thunberg, 1815) | |
Synonyms [1] | |
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Dociostaurus maroccanus, commonly known as the Moroccan locust , is a grasshopper in the insect family Acrididae. It is found in northern Africa, southern and eastern Europe and western Asia. It lives a solitary existence but in some years its numbers increase sharply, and it becomes gregarious and congregates to form swarms which can cause devastation in agricultural areas. The species was first described by Carl Peter Thunberg in 1815.
The range of the Moroccan locust extends from the Canary Islands and Madeira in the west to Kazakhstan and Afghanistan in the east. In Africa it is found in Algeria, Egypt, Libya, Morocco and Tunisia. In Europe it is found in France, Portugal, Spain, Italy and the Balkan peninsula. It is also found in the Middle East and Central Asia. [2]
The eggs are laid in pods with about thirty eggs each. The nymphs ("hoppers") resemble wingless adults. They moult five times, each instar having larger wing pads. The adult female locust is 20 to 38 millimetres long and the male 16 to 28 millimetres long. The body colour is yellowish gray with dark patches. There is a creamy coloured cross-shape on the prothorax. The elytra are large and transparent, sometimes speckled with brown, and the wings are colourless and strongly veined. The hind legs are powerful, the femur often being banded with black while the tibia is usually red. The time taken from hatching to maturity is about thirty days. [2]
The Moroccan locust lives in arid areas between 500 and 1000 metres above sea level. It breeds in undisturbed soil and does not reproduce in arable fields. [2] The optimum rainfall for development in the months from March to May is one hundred millimetres. If the spring is unusually hot and precipitation is low, and especially if it is the second dry spring in succession, numbers of young locusts can increase dramatically. Mass outbreaks of nymphs occur and destroy the vegetation where they hatch. From the second instar stage onwards, the hoppers aggregate and the population density can reach several thousand nymphs per square metre. [3] The hoppers move downhill to fertile regions damaging crops as they go. Multiple generations can be present, and the numbers of adults can build up rapidly, reaching 300 individuals per square metre in extreme cases. Winged adults take flight en masse at low altitudes, travelling at eight to ten metres per second and may migrate sixty kilometres during the season. Numbers begin to diminish in June when the vegetation is also declining. [2]
This locust has traditionally been considered a serious agricultural threat. Its large numbers, voracious appetite, ability to eat almost all forms of vegetation and ability to migrate in swarms make it a feared pest. However, outbreaks of swarming activity have diminished in recent years and in some areas the Moroccan locust has become rare. This is particularly noticeable in European countries where the species no longer seems to be an economic threat while in north Africa and parts of central Asia it still flourishes. These alterations in population dynamics may be due to a change in climatic conditions with a decrease in spring rainfall. A more important factor may be the increase of cropland at the expense of grassland, since the female needs undisturbed soil in which to deposit her eggs. At the other extreme, overgrazing and deforestation in some areas provide a favourable environment for the species to thrive and remain destructive. [4]
In a study in Iran in 1956 it was found that outbreaks of locusts originated from just eight distinct areas. All these districts were in mountain foothills that had been forested or in other rolling foothills. In all cases the vegetation consisted of annual grasses or shrub steppe communities with annual grasses and dwarf shrubs. There was a good supply of green food in winter which increased the longevity and fertility of the females. [5] Outside these districts, the Moroccan locust had solitary populations whereas inside them, in some years, the locust became gregarious. [6] The locust needs firm, bare soil for egg laying and when they hatch, the nymphs move to progressively taller and denser vegetation. [5] These districts principally had rainfall in winter and early spring, averaging 250 to 350 millimetres. The temperature averaged about 17 °C in the months immediately before and after egg laying. The sites were heavily grazed by sheep and goats, especially in winter. The conclusion of the study was that quite specific climate, vegetation, and topographical conditions were needed for breeding to take place. [6]
Locusts are a group of certain species of short-horned grasshoppers in the family Acrididae that have a swarming phase. These insects are usually solitary, but under certain circumstances they become more abundant and change their behaviour and habits, becoming gregarious. No taxonomic distinction is made between locust and grasshopper species; the basis for the definition is whether a species forms swarms under intermittently suitable conditions.
Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago.
The desert locust is a species of locust, a periodically swarming, short-horned grasshopper in the family Acrididae. They are found primarily in the deserts and dry areas of northern and eastern Africa, Arabia, and southwest Asia. During population surge years, they may extend north into parts of western Spain and southern Italy, south into Eastern Africa, and east in northern India. The desert locust shows periodic changes in its body form and can change in response to environmental conditions, over several generations, from a solitary, shorter-winged, highly fecund, non-migratory form to a gregarious, long-winged, and migratory phase in which they may travel long distances into new areas. In some years, they may thus form locust plagues, invading new areas, where they may consume all vegetation including crops, and at other times, they may live unnoticed in small numbers.
The migratory locust is the most widespread locust species, and the only species in the genus Locusta. It occurs throughout Africa, Asia, Australia and New Zealand. It used to be common in Europe but has now become rare there. Because of the vast geographic area it occupies, which comprises many different ecological zones, numerous subspecies have been described. However, not all experts agree on the validity of some of these subspecies.
The red locust is a large grasshopper species found in sub-Saharan Africa. Its name refers to the colour of its hind wings. It is sometimes called the criquet nomade in French, due to its nomadic movements in the dry season. When it forms swarms, it is described as a locust.
The Australian plague locust is a native Australian insect in the family Acrididae, and a significant agricultural pest.
The Senegalese grasshopper is a medium-sized grasshopper species found in the Sahel region of Africa, the Canary Islands, Cape Verde Islands, and West Asia. Although not called a locust in English, this species shows gregarious behaviour and some morphological change on crowding. In many parts of the Sahel, this species may cause greater year-on-year crop damage than better-known locusts, attacking crops such as the pearl millet.
The brown locust is a medium-sized small locust species in the monotypic genus Locustana. It is found in Southern Africa and shows classic gregarious behaviour with phase polymorphism on crowding.
Chelisoches morio, also known as the black earwig, are part of the family Chelisochidae. It is an omnivore that can be found worldwide, however it is most prominent in tropical areas, Pacific islands, the Pacific Northwest, and damp environments. The adults are jet black and can range in size from 18 to 25mm in size, though some have grown to be 36mm. The males cerci are widely separated and serrated compared to the female. The forceps are used for prey capture, defense, fighting and courtship.
Eurygaster integriceps is a species of shield bug in the family Scutelleridae, commonly known as the sunn pest or corn bug. It is native to much of northern Africa, the Balkans and western and central Asia. It is a major pest of cereal crops especially wheat, barley and oats.
Archipsocus nomas is a web-spinning barklouse, a psocid in the insect family Archipsocidae. It is found in the southeast of the United States, living gregariously on trees, feeding on and lichen and fungi and spinning a web that adheres to the trunk and large branches in sheets. The webs are thought to protect the barklice from predators and neither the insects nor the webs cause damage to the trees.
Calliptamus italicus, the Italian locust, is a species of 'short-horned grasshopper' belonging to the family Acrididae, subfamily Calliptaminae.
Schistocerca americana is a species of grasshopper in the family Acrididae known commonly as the American grasshopper and American bird grasshopper. It is native to North America, where it occurs in the eastern United States, Mexico, and the Bahamas. Occasional, localized outbreaks of this grasshopper occur, and it is often referred to as a locust, though it lacks the true swarming form of its congener, the desert locust.
Patanga succincta, the Bombay locust, is a species of locust found in India and southeast Asia. Usually a solitary insect, only in India has it has exhibited swarming behaviour. The last plague of this locust was in that country between 1901 and 1908 and there have not been any swarms since 1927. It is thought that the behaviour of the insects has altered because of changing practices in agricultural land use.
Valanga nigricornis, the Javanese grasshopper, is a species of grasshopper in the subfamily Cyrtacanthacridinae of the family Acrididae. It is found in southeastern Asia, the type location being Singapore. It was first described by the German zoologist Hermann Burmeister in 1838. There are more than twenty subspecies, most of which are endemic to different island groups; the subspecies V. nigricornis nigricornis is the type for the genus Valanga.
Locusta migratoria manilensis, commonly known as the Oriental migratory locust, is a subspecies of the migratory locust in the family Acrididae. It is sufficiently different in size and structure from the African migratory locust to be considered a distinct subspecies of the migratory locust. It is found in southeastern Asia and is an important agricultural pest in the region. It is normally a solitary insect but when conditions are suitable, it enters into a gregarious phase when the young form into bands which move together and the adults into swarms. Although outbreaks may have recently been fewer in number and size because of changes in agricultural practices and better locust detection, the insects remain active as crop pests and the potential for outbreaks is still present.
Locusta migratoria migratorioides, commonly known as the African migratory locust, is a subspecies of the migratory locust family Acrididae.
Valanga irregularis belongs to the family Acrididae. The distribution is restricted in the Australian tropics and subtropics. The species is the largest grasshopper of the continent. Usually the animal lives a solitary mode of life.
Phymateus aegrotus, sometimes called the blue bush locust or East African bush locust, is a pest species of grasshopper in the family Pyrgomorphidae. Unlike "locusts" the adults are not known to change their morphology on crowding, but at the hopper stage, marching behaviour of small bands may occur.
2019–2022 locust infestation is a pest outbreak of desert locusts which is threatening the food supply across the regions of East Africa, the Arabian Peninsula, and the Indian subcontinent. The outbreak is the worst in 70 years in Kenya and the worst in 25 years in Ethiopia, Somalia and India. The plague began in June 2019 and continued into 2020, although locust swarms have experienced steady declines in population and geographical reach from May to October, and as of November 2020, primarily found in the Horn of Africa and Yemen.