| "Eolouka" | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Clade: | Diphoda |
| Clade: | Discoba |
| Phylum: | Eolouka Cavalier-Smith, 2013 [1] stat. nov. [2] |
| Groups included | |
| Cladistically included but traditionally excluded taxa | |
Eolouka is a paraphyletic phylum of protists localized in the clade Discoba. It contains two lineages: Jakobea and Tsukubea, the last containing only one genus, Tsukubamonas . [1] [2]
In 1999 Cavalier-Smith proposed a new paraphyletic phylum of flagellates called Loukozoa, containing only the jakobids. [3] In 2013 it was modified to contain three subphyla: [1]
However, in later years these three groups were raised to the rank of phylum. [4] [2]
The alveolates are a group of protists, considered a major clade and superphylum within Eukarya. They are currently grouped with the stramenopiles and Rhizaria among the protists with tubulocristate mitochondria into the SAR supergroup.
The axodines are a group of unicellular stramenopiles that includes silicoflagellate and rhizochromulinid algae, actinomonad heterotrophic flagellates and actinophryid heliozoa. Alternative classifications treat the dictyochophytes as heterokont algae, or as Chrysophyceae. Other overlapping taxonomic concepts include the Actinochrysophyceae, Actinochrysea or Dictyochophyceae sensu lato. The grouping was proposed on the basis of ultrastructural similarities, and is consistent with subsequent molecular comparisons.
Chromista is a proposed but polyphyletic biological kingdom, refined from the Chromalveolata, consisting of single-celled and multicellular eukaryotic species that share similar features in their photosynthetic organelles (plastids). It includes all eukaryotes whose plastids contain chlorophyll c and are surrounded by four membranes. If the ancestor already possessed chloroplasts derived by endosymbiosis from red algae, all non-photosynthetic Chromista have secondarily lost the ability to photosynthesise. Its members might have arisen independently as separate evolutionary groups from the last eukaryotic common ancestor.
Thomas (Tom) Cavalier-Smith, FRS, FRSC, NERC Professorial Fellow, was a professor of evolutionary biology in the Department of Zoology, at the University of Oxford.
The metamonads are a large group of flagellate amitochondriate microscopic eukaryotes. Their composition is not entirely settled, but they include the retortamonads, diplomonads, and possibly the parabasalids and oxymonads as well. These four groups are all anaerobic, occurring mostly as symbiotes or parasites of animals, as is the case with Giardia lamblia which causes diarrhea in mammals.
Amorphea is a taxonomic supergroup that includes the basal Amoebozoa and Obazoa. That latter contains the Opisthokonta, which includes the Fungi, Animals and the Choanomonada, or Choanoflagellates. The taxonomic affinities of the members of this clade were originally described and proposed by Thomas Cavalier-Smith in 2002.
Loukozoa is a proposed taxon used in some classifications of eukaryotes, consisting of the Metamonada and Malawimonadea. Ancyromonads are closely related to this group, as sister of the entire group, or as sister of the Metamonada. Amorphea may have emerged in this grouping, specifically as sister of the Malawimonads.
Bigyra is a phylum of microscopic eukaryotes that are found at the base of the Stramenopiles clade. It includes three well-known heterotrophic groups Bicosoecida, Opalinata and Labyrinthulomycetes, as well as several small clades initially discovered through environmental DNA samples: Nanomonadea, Placididea, Opalomonadea and Eogyrea. The classification of Bigyra has changed several times since its origin, and its monophyly remains unresolved.
Opalozoa is a subphylum of heterotrophic protists of the phylum Bigyra, and is the sister group to Sagenista. Opalozoans are non-photosynthetic heterokonts that are ancestrally phagotrophic but many times have evolved to be osmotrophic saprotrophs in the gut of vertebrate animals.
Halvaria is a taxonomic grouping of protists that includes Alveolata and Stramenopiles (Heterokonta).
The initial version of a classification system of life by British zoologist Thomas Cavalier-Smith appeared in 1978. This initial system continued to be modified in subsequent versions that were published until he died in 2021. As with classifications of others, such as Carl Linnaeus, Ernst Haeckel, Robert Whittaker, and Carl Woese, Cavalier-Smith's classification attempts to incorporate the latest developments in taxonomy., Cavalier-Smith used his classifications to convey his opinions about the evolutionary relationships among various organisms, principally microbial. His classifications complemented his ideas communicated in scientific publications, talks, and diagrams. Different iterations might have a wider or narrow scope, include different groupings, provide greater or lesser detail, and place groups in different arrangements as his thinking changed. His classifications has been a major influence in the modern taxonomy, particularly of protists.
Platysulcus is an eukaryotic microorganism that was recently discovered to be the earliest diverging lineage of the Heterokont phylogenetic tree. It is the only member of the family Platysulcidae, order Platysulcida and class Platysulcea, of uncertain taxonomic position within the phylum Bigyra. It contains the only species P. tardus.
The Orthokaryotes are a proposed Eukaryote clade consisting of the Jakobea and the Neokaryotes. Together with its sister Discicristata it forms a basal Eukaryote clade. They are characterized by stacked Golgi, orthogonal centrioles, and two opposite posterior ciliary roots.
The neokaryotes are a proposed eukaryote clade consisting of the unikonts and the bikonts as sister of for instance the Jakobea. It arises because the Euglenozoa, Percolozoa, Tsukubea, and Jakobea are seen in this view as more basal eukaryotes. These four groups, are traditionally grouped together in the Discoba. However, the Discoba may well be paraphyletic as the neokaryotes may have emerged in them.
The Scotokaryotes (Cavalier-Smith) is a proposed basal Neokaryote clade as sister of the Diaphoretickes. Basal Scotokaryote groupings are the Metamonads, the Malawimonas and the Podiata. In this phylogeny the Discoba are sometimes seen as paraphyletic and basal Eukaryotes.
The cortical alveolum is a cellular organelle consisting of a vesicle located under the cytoplasmic membrane, to which they give support. They have been defined as membrane sacs that strengthen the cellular cortex through the firm fixation to the underlying membrane and microtubules. They typically form a continuous layer that acts as a flexible film, although they can also constitute a semi-rigid structure or the scales of a theca.
Gyrista is a phylum of heterokont protists containing three diverse groups: the mostly photosynthetic Ochrophyta, the parasitic Pseudofungi, and the recently described group of nanoflagellates known as Bigyromonada. Members of this phylum are characterized by the presence of a helix or a double helix/ring system in the ciliary transition region.
Malawimonads are a small group of microorganisms with a basal position in the evolutionary tree of eukaryotes, containing only three recognized species. They're considered part of a possibly paraphyletic group known as "Excavata".
Commation cryoporinum is a species of heterotrophic protists discovered in 1993 in Antarctic waters. It is one of two species in the Commatiida, an order of stramenopiles closely related to actinophryids, a group of heliozoan protists, and to raphidophytes, a group of algae.
Raphopoda is a grouping of heterotrophic protists. It contains the heterotroph organisms within class Raphidomonadea, classified as two orders: Commatiida, comprised by the sole genus of flagellates Commation, and Actinophryida, an order of heliozoa, amoebae with stiff specialized pseudopodia called axopodia.
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