Chondrichthyes is a class of jawed fish that contains the cartilaginous fish or chondrichthyians, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fish, which have skeletons primarily composed of bone tissue. Chondrichthyes are aquatic vertebrates with paired fins, paired nares, placoid scales, conus arteriosus in the heart, and a lack of opecula and swim bladders. Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates.
Conodonts are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which are usually found in isolation and are now called conodont elements. Knowledge about soft tissues remains limited. They existed in the world's oceans for over 300 million years, from the Cambrian to the beginning of the Jurassic. Conodont elements are widely used as index fossils, fossils used to define and identify geological periods.
Agnatha is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both living (cyclostomes) and extinct species. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes.
Gnathostomata are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. In addition to opposing jaws, living gnathostomes have true teeth, paired appendages, the elastomeric protein of elastin, and a horizontal semicircular canal of the inner ear, along with physiological and cellular anatomical characters such as the myelin sheaths of neurons, and an adaptive immune system that has the discrete lymphoid organs of spleen and thymus, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene.
The Amiiformes order of fish has only two extant species, the bowfins: Amia calva and Amia ocellicauda, the latter recognized as a separate species in 2022. These Amiiformes are found in the freshwater systems of North America, in the United States and parts of southern Canada. They live in freshwater streams, rivers, and swamps. The order first appeared in the Triassic, and the extinct members include both marine and freshwater species, many of which are morphologically disparate from bowfins, such as the caturids.
Arandaspida is a taxon of very early, jawless prehistoric fish which lived during the Ordovician period. Arandaspids represent the oldest known craniates, a proposed group of chordates that contain all chordates with a cartilage-derived skull, and hagfish. The group represents a subclass within the class Pteraspidomorphi, and contains only one order, the Arandaspidiformes. The oldest known genus of this group is Sacabambaspis found in South America.
Heterostraci is an extinct subclass of pteraspidomorph, Ostracoderm, jawless vertebrate that lived primarily in marine and estuary environments. Heterostraci existed from the mid-Ordovician to the conclusion of the Devonian.
Protaspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protaspidids were once thought to represent a transitional form between the Pteraspididae and the Psammosteida, bearing the broad head shield shape of the latter, due to a more benthic (bottom-dwelling) existence, but recent phylogenical comparisons demonstrate that the protaspidids are actually highly derived pteraspidids, and that the anchipteraspidids, the most primitive of pteraspidids, are the sister-group of the Psammosteids.
Thelodonti is a class of extinct Palaeozoic jawless fishes with distinctive scales instead of large plates of armor.
Anaspida is an extinct group of jawless fish that existed from the early Silurian period to the late Devonian period. They were classically regarded as the ancestors of lampreys, but it is denied in recent phylogenetic analysis, although some analysis show these group would be at least related. Anaspids were small marine fish that lacked a heavy bony shield and paired fins, but were distinctively hypocercal.
Pteraspidomorphi is an extinct class of early jawless fish. They have long been regarded as closely related or even ancestral to jawed vertebrates, but the few characteristics they share with the latter are now considered as basal traits for all vertebrates.
Arandaspis prionotolepis is an extinct species of jawless fish that lived in the Ordovician period, about 480 to 470 million years ago. Its remains were found in the Stairway Sandstone near Alice Springs, Australia in 1959, but it was not determined that they were the oldest known vertebrates until the late 1960s. Arandaspis is named after a local Indigenous Australian people, the Aranda.
Drepanaspis is an extinct genus of primitive jawless Ostracoderms from the Early Devonian that belonged to the phylum Chordata, infraphylum Agnatha, class Pteraspidomorphi, and the subclass Heterostraci. Drepanaspis are assumed to have lived primarily in marine environments and is most commonly characterized by their ray-like, heavily armored bodies, along with their lack of paired fins and jaws.
Astraspida, or astraspids, are a small group of extinct armored jawless vertebrates, which lived in the Middle Ordovician in North America. They are placed among the Pteraspidomorphi because of the large dorsal and ventral shield of their head armor. They are represented by a single genus, Astraspis, including possibly two species, A. desiderata and A. splendens but their remains are fairly abundant in Ordovician sandstones of the USA and Canada (Quebec). The head armor of Astraspis is rather massive, with a series of ten gill openings lining the margin of the dorsal shield, and laterally placed eyes. The dorsal shield is ribbed by strong longitudinal crests, and the tail is covered with large, diamond-shaped scales. They are often grouped together with the Arandaspidida.
Albulidae is a family of fish, commonly known as the bonefishes, that are popular as game fish in Florida, select locations in the South Pacific and the Bahamas and elsewhere. The family is small, with 11 species in 3 genera. Presently, the bonefishes are in their own order: Albuliformes. The families Halosauridae and Notacanthidae were previously classified in this order, but are now, according to FishBase, given their own order Notacanthiformes. The largest bonefish caught in the Western Hemisphere is a 16-pound, 3 ounce example caught off Islamorada, Florida, on March 19, 2007.
Sacabambaspis is an extinct genus of jawless fish that lived in the Ordovician period. Sacabambaspis lived in shallow waters on the continental margins of Gondwana. It is the best known arandaspid with many specimens known.
Porolepiformes is an order of prehistoric lobe-finned fish which lived during the Devonian period. They are thought to represent the sister group to lungfish. The group contains two families: Holoptychiidae and Porolepididae.
Tarasiiformes is an extinct order of prehistoric ray-finned fish.
Birkeniiformes is an extinct order of jawless fish belonging to the class Anaspida.
Amia, commonly called bowfin, is a genus of bony fish related to gars in the infraclass Holostei. They are regarded as taxonomic relicts, being the sole surviving species of the order Amiiformes, which dates from the Jurassic to the Eocene, persisting to the present. There are two living species in Amia, Amia calva and Amia ocellicauda, and a number of extinct species which have been described from the fossil record.
