Geophilus carpophagus | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Myriapoda |
Class: | Chilopoda |
Order: | Geophilomorpha |
Family: | Geophilidae |
Genus: | Geophilus |
Species: | G. carpophagus |
Binomial name | |
Geophilus carpophagus Leach, 1815 | |
Geophilus carpophagus is a species of soil centipede in the family Geophilidae, widely considered to be a type species of the genus Geophilus (along with G. electricus ). [1] It grows up to 60 millimeters in length, with an orange/tan body bearing a distinctive purplish marbled pattern (also seen in Henia vesuviana ). [2] Males of this species have 51 to 55 pairs of legs; females have 53 to 57. [3]
Recent studies indicate that G. carpophagus is actually a group of closely related species (dubbed the carpophagus species-complex). So far only three species have been clearly defined; G. carpophagus (Leach, 1814), G. easoni (Arthur, et al., 2001) from Europe, and G. arenarius (Meinert, 1870) from North Africa. [3]
Geophilus carpophagus is most commonly found across southeast England, though specimens have also been found in mainland Europe, North Africa, and the Canarie Islands, with a viable population discovered in Finland in 2018. [4] In the north of England, it becomes entirely coastal, where it can be found in cliff sites above the high tide mark. Away from the coast, it is usually found living 1 meter or more above the ground in rocks, walls, buildings, and trees. G. carpophagus seemingly has stronger adhesive abilities than G. easoni, presumably to help with climbing. G. carpophagus nests in drier sites than most other British geophilomorphans, which may be connected to its reduced size and number of coxal pores compared to G. easoni. [5]
Geophilus carpophagus has a less aggressive defense response than G. easoni (recoiling instead of rearing and bearing poison claws). In laboratory cultures, mothers that were deliberately disturbed and left their brood would return to coil around it again, a behavior that has never been observed in any other geophilomorph species. [5]
Geophilus flavus is a terrestrial, soil-dwelling, species of centipede in the Geophilidae family. G. flavus occurs in a range of habitats across central Europe, North America, Australia and other tropical regions. Geophilomorph centipedes, like centipedes generally, are primary predators, hunting predominantly in underground soil burrows or above ground leaf litter. Their consumption behaviours are influenced by environment and seasonal factors. Given their lack of economic value and marginal medical significance, G.flavus remains largely understudied in mainstream research. Some recent studies have detailed the evolutionary development of G.flavus and Geophilidae generally, illustrating developed predatory features like forcipule venom glands.
Pachymerium ferrugineum is a species of centipede in the family Geophilidae.
The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae.
Geophilus hadesi is a species of soil centipede in the family Geophilidae, named after Hades, king of the underworld in Greek mythology. This species is found in the caves of the Velebit Mountains of Croatia and characterized by relatively elongated trunk segments and appendages, including unusually long claws of the legs. These centipedes have 33 pairs of legs and can be as long as 28 mm. Like other geophilomorhpans, this species lacks sight, has a flattened trunk, and is well adapted to underground life. Along with Geophilus persephones it is one of the only two known troglomorphic geophilomorphs and can even be found in Lukina jama, the 15th deepest cave in the world.
Mecistocephalidae is a monophyletic family of centipedes in the order Geophilomorpha. It is the only family in the suborder Placodesmata. Most species in this family live in tropical or subtropical regions, but some occur in temperate regions. This family is the third most diverse in the order Geophiliomorpha, with about 170 species, including about 130 species in the genus Mecistocephalus.
Geophilus is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, anterior porefields, complete or nearly complete coxo-pleural sutures at the prosternum, and incomplete chitin-lines. Centipedes in this genus range from 1 cm to 8 cm in length. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus.
Geophilus persephones is a species of soil centipede in the family Geophilidae discovered in 1999. This species is named after Persephone, the queen of the underworld in Greek mythology, and found in caves in the Gouffre de la Pierre Saint-Martin. It has elongated antennae and legs as well as abundant sensory setae, and like other geophilomorhps it lacks sight, has a flattened trunk, and is well adapted to underground life. This species was the first troglomorphic geophilomorph ever discovered and one of the only two in existence along with Geophilus hadesi. Known from a single male specimen, this species has only 29 pairs of legs, one of only two species in the Geophilidae family to have so few leg pairs.
Geophilus electricus is a species of soil centipede in the family Geophilidae found across temperate Europe, especially Britain and Ireland. Originally discovered and named Scolopendra electrica in 1758, it was later moved to its own genus, Geophilus. As the first species described in this genus, it is considered by some to be the type species. It is yellowish-orange in color and can grow up to 45 millimeters, with 55 to 75 leg pairs, a distinct carpophagus fossae on the anterior sternites, and a distinctive arrangement of the coxal pores of the last leg pair. Like many geophilomorphans, G. electricus has been known to glow in the dark, most likely as a defensive maneuver.
Geophilus easoni is a species of soil centipede in the family Geophilidae found throughout Britain and Ireland, though its range extends through western France to at least the foothills of the Pyrenees. Until 2001, it was considered synonymous with G. carpophagus, and most pre-2001 records probably refer to G. easoni. It's typically shorter than G. carpophagus, with fewer leg pairs, uniform tan/chestnut coloring, and a greater size and number of coxal pores, as well as a darkly pigmented mid-piece of the labrum which bears blunt teeth. Specimens in north-western Iberia typically have a wider range of leg pairs. It nests on the ground surface under stones and dead wood.
Geophilus arenarius is a species of soil centipede in the family Geophilidae found in northwest Africa, specifically near Annaba, Algeria. This species is frequently misidentified with G. electricus, and as part of the carpophagus species-complex, is closely related to both G. carpophagus and G. easoni, though it differs mainly by lacking a transverse suture on the head and peculiar integumental features (carpophagus-structures) along the trunk, as well as having relatively stouter antennae and forcipular coxosternite. G. arenarius is distinctly smaller at full growth than G. carpophagus, with usually blunter and more sclerotised tubercles lining the intermediate part of the labrum and a minute denticle at the basis of the forcipular tarsungula. It has fewer bristles lining the lateral parts of the labrum than G. easoni as well as a generally higher number of legs and a more greyish coloured trunk. An examination of 36 G. arenarius syntypes indicates that males of this species have 55 pairs of legs, whereas females have 55 to 59 leg pairs, with 57 as the most common number.
Geophilus proximus is a species of soil centipede in the family Geophilidae found in the northern part of the Palearctic and widespread across the entire Baltic basin, though it reaches as far as the Arctic Circle and has been introduced through human agency to northern, central, and eastern parts of Kazakhstan. It was recorded once with certainty in Britain from Unst in the Shetland Islands; distribution in the rest of Europe is difficult to assess because of frequent misidentifications of the species. Populations from northern Europe are mostly parthenogenetic.
Geophilus impressus is a species of soil centipede in the family Geophilidae found all over Europe, and has also been recorded in North Africa. It lives frequently in endogean habitats; in Sardinia it's found mostly in Quercus ilex woods, but also in Mediterranean shrub, open habitats, and maquis. It lives anywhere from sea level to 1700 meters above it, sometimes in caves.
Geophilus gracilis is a species of soil centipede in the family Geophilidae found most commonly in Britain and Ireland, though specimens have also been recorded in Chile, France, Greece, and Algeria. It lives under mud and stones along the coast near or below the high tide mark, grows up to 30 millimeters in length, and is bright yellow in color with a darker reddish head. Males of this species have 51 to 57 pairs of legs; females have 51 to 61 leg pairs. This species is often confused with G. flavus and, in coastal locations, G. osquidatum.
Geophilus osquidatum is a species of soil centipede in the family Geophilidae found in western Europe, from mainland Spain through western France to Britain and Ireland, though it's also been recorded in Italy, Czech Republic, and Germany. Males of this species have 51 to 57 pairs of legs; females have 53 to 63 leg pairs. This species grows up to 30 millimeters and is bright yellow with a darker reddish head. Because of this, it's often confused with G. flavus and G. gracilis. Its subspecies, G. osquidatum porosum, was found synonymous with G. flavus. In Britain, G. osquidatum is found in a wide range of habitats including woodland, grassland, and coastal shingle as well as gardens and waste ground.
Geophilus vittatus, also known as the diamondback soil centipede, is a species of soil centipede in the family Geophilidae widespread in North America and named for the dark band of diamond-shaped markings across its back. G. vittatus grows up to 52 millimeters in length, though it averages 25–40, and ranges from brown- to orange-yellow in color, with 49–53 leg pairs in females, 49–51 leg pairs in males, and a lack of consolidated carpophagus structures. It can be found under any debris on the forest floor, but mostly under loose bark and occasionally near the sea.
Geophilus varians is a species of soil centipede in the family Geophilidae found in North America, particularly from South Carolina to Indiana, Ohio, Pennsylvania, and Virginia. It grows up to 40 millimeters, though it averages 30-35, ranges in color from light faded orange to yellow or whitish yellow with a deeper and brighter head, and has 53-59 leg pairs in males and 55-61 in females, as well as a complete lack of consolidated paxilli and sacculi, concealed prebasal plate, and unusually long ultimate legs.
Geophilus alzonis is a species of soil centipede in the family Geophilidae found in Monte Alzo near Tolosa in Spain, which it was named after. It's a poorly defined species that was described as being light yellow with a light chestnut brown head and 13–14 mm long, with small but numerous leg bristles, sternites bearing three longitudinal furrows, a carpophagus formation from segments 3–6 to segments 11th-13, and 37-51 leg pairs.
Geophilus truncorum is a species of soil centipede in the family Geophilidae found across Western Europe, though it reaches as far as Poland, Italy, and Morocco. This centipede is relatively small, growing up to 20mm in length, with a yellow or orangeish brown body and dark yellow or brown head, denser and shorter hair than most Geophilus species, a main plate almost as elongated as in G. flavus (115:100), and distinct carpophagus fossae on the anterior sternites. Males of this species have 35 to 41 pairs of legs; females have 37 to 41.
Geophilus bosniensis is a species of soil centipede in the family Geophilidae endemic to Bosnia and Herzegovina. It grows up to 30 millimeters and has 75 leg pairs, as well as sternites unseparated in the median but with a suture line, and sternal pore areas in the trunk segments only. Overall, the identity and phyletic position of this centipede are uncertain.
The centipedes or Chilopoda are divided into five orders, which are grouped into two subclasses, Pleurostigmomorpha and Notostigmomorpha, the latter of which comprises only one order, the Scutigeromorpha.
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