Gravicalymene

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Gravicalymene arcuata Price, 1982, Rawtheyan Stage, Dolhir Formation, Cynwyd Forest, near Corwen, Wales. Gravicalymene arcuata.jpg
Gravicalymene arcuata Price, 1982, Rawtheyan Stage, Dolhir Formation, Cynwyd Forest, near Corwen, Wales.

Gravicalymene
Gravicalymene yamakoshii - National Museum of Nature and Science, Tokyo - DSC07017.JPG
Gravicalymene yamakoshii Kobayashi & Hamada, 1977. From the Devonian of Japan.
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Phacopida
Family: Calymenidae
Genus: Gravicalymene
Species

See text.

Gravicalymene Shirley, 1936, [1] is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and family Calymenidae. Species included in this genus have previously been allocated to Calymene Brongniart 1822, [2] Flexicalymene Shirley, 1936. and Sthenarocalymene Siveter 1977. [3]

Contents

Rarest within the genus is the Middle Ordovician species Gravicalymene magnotuberculata, which is also amongst the rarest of all Calymenidae and regionally confined to two exposures in New York State. G. magnotuberculata is noted for its extremely pustulose exoskeleton, bell-shaped glabella and lack of complete articulated specimens.

Type species

Other species

Some known species and locations include:

Related Research Articles

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.

Silurian Third period of the Paleozoic Era 444-419 million years ago

The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.

Trilobite Class of extinct, Paleozoic arthropods

Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 20,000 species having been described.

John William Salter

John William Salter was an English naturalist, geologist, and palaeontologist.

In geology, the Llandeilo Group is the middle subdivision of the British Ordovician rocks. It was first described and named by Sir Roderick Murchison from the neighborhood of Llandeilo in Carmarthenshire. In the type area it consists of a series of slaty rocks, shales, calcareous flagstones and sandstones; the calcareous middle portion is sometimes termed the Llandeilo limestone; and in the upper portion volcanic rocks are intercalated.

<i>Calymene</i>

Calymene Brongniart, 1822, is a genus of trilobites in the order Phacopida, suborder Calymenina, that are found throughout North America, North Africa, and Europe in primarily Silurian outcrops. Calymene is closely related to Flexicalymene, and both genera are frequently found enrolled. Calymene trilobites are small, typically 2 cm in length. The cephalon is the widest part of the animal and the thorax usually has 13 segments.

<i>Flexicalymene</i> Genus of trilobites (fossil)

Flexicalymene Shirley, 1936. is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and Family Calymenidae. Flexicalymene specimens can be mistaken for Calymene, Gravicalymene, Diacalymene and a few other Calymenina genera. They are used as an index fossil in the Ordovician. Ohio and North America are particularly known for being rich with Flexicalymene fossils.

<i>Kainops</i> Genus of trilobites

Kainops is a genus of trilobites from the family Phacopidae, order Phacopida. It can be distinguished from Paciphacops by the greater number of facets to the eye. The form of the furrow between the palpebral area and the palpebral lobe also distinguishes Kainops from the genera Paciphacops and Viaphacops.

<i>Calymene blumenbachii</i>

Calymene blumenbachii Brongniart in Desmarest (1817), sometimes erroneously spelled blumenbachi, is a species of trilobite discovered in the limestone quarries of the Wren's Nest in Dudley, England. Nicknamed the Dudley Bug or Dudley Locust by 18th-century quarrymen it became a symbol of the town and featured on the Dudley County Borough Council coat-of-arms. Calymene blumenbachii is commonly found in Silurian rocks and is thought to have lived in the shallow waters of the Silurian, in low-energy reefs. This particular species of Calymene is unique to the Wenlock series in England, and comes from the Wenlock Limestone Formation in Much Wenlock and the Wren's Nest in Dudley. These sites seem to yield trilobites more readily than any other areas on the Wenlock Edge, and the rock here is dark grey as opposed to yellowish or whitish as it appears on other parts of the Edge, just a few miles away, in Church Stretton and elsewhere. This suggests local changes in the environment in which the rock was deposited.

<i>Erbenochile</i>

Erbenochile is a genus of spinose phacopid trilobite, of the family Acastidae, found in Lower to Middle Devonian age rocks from Algeria and Morocco. Originally described from an isolated pygidium, the first complete articulated specimen of E. erbeni revealed the presence of extraordinarily tall eyes:

"Straight-sided towers of lenses... with [up to] 18 lenses in a vertical file"

<i>Balizoma</i> Genus of trilobites

Balizoma is a genus of trilobites from the family Encrinuridae established by David J. Holloway in 1980. It has only been found in rocks of Silurian age. Its type species, B. variolaris, is currently the only named species of the genus, and is found in England. The neotype of B. variolaris was collected from the Much Wenlock Limestone Formation at Dudley, West Midlands. That specimen was first illustrated in Sir Roderick Impey Murchison's classic book, The Silurian System. B. variolaris was the original "strawberry-headed" trilobite of Dudley, so-named because of its nodular glabellar tubercles, and well known to early trilobite collectors. Additional species were originally assigned to Balizoma, but were subsequently placed in other encrinurine genera.

<i>Aulacopleura</i> Extinct genus of trilobites

Aulacopleura is a genus of proetid trilobite that lived from the Middle Ordovician to the Middle Devonian. Some authors may classify this group as subgenus Otarion (Aulacopleura). The cephalon is semicircular or semielliptical, with border and preglabellar field. The glabella is short, with or without defined eye ridges connecting it with eyes of variable size. Spines at the rear outer corners of the cephalon are present, typically reaching back to the 2nd to 4th thorax segment. The 'palate' is not connected to the dorsal shield of the cephalon. The cephalon is pitted, or has small tubercles. The thorax has up to 22 segments. The pleural ends are usually rounded. The pygidium is small (micropygous), with an even margin. A. koninckii had a modern type of compound eye.

Cynwyd Forest Quarry

Sometimes referred to in literature as 'Bwlch y Gaseg' and in very close proximity to the area named as such on OS Map 1888-1913, the Cynwyd Forest Quarry is a Site of Special Scientific Interest (SSSI) located aside a track within the Cynwyd Forest near Corwen, Denbighshire, North Wales. It was described by Rushton et al. (2000) and exposes Late Ordovician micaceous siltstones and mudstones of the Dolhir Formation which yields a rich shelly (brachiopod) fauna. Trilobites are represented by several genera although the fauna is dominated by Gravicalymene arcuata Price, 1982. Bivalves, bryozoans and various Echinodermata are also present. Examples of most of the fossils listed below are illustrated and briefly described in "Fossils of the Upper Ordovician" by Harper and Owen (Eds.).

Jeffersonville Limestone

The Devonian Jeffersonville Limestone is a mapped bedrock unit in Indiana and Kentucky. It is highly fossiliferous.

Paleontology in Wisconsin

Paleontology in Wisconsin refers to paleontological research occurring within or conducted by people from the U.S. state of Wisconsin. The state has fossils from the Precambrian, much of the Paleozoic, and the later part of the Cenozoic. Most of the Paleozoic rocks are marine in origin. Because of the thick blanket of Pleistocene glacial sediment that covers the rock strata in most of the state, Wisconsin’s fossil record is relatively sparse. In spite of this, certain Wisconsin paleontological occurrences provide exceptional insights concerning the history and diversity of life on Earth.

<i>Trinodus</i> Extinct genus of trilobites

Trinodus is a very small to small blind trilobite, a well known group of extinct marine arthropods, which lived during the Ordovician, in what are now the Yukon Territories, Virginia, Italy, Czech Republic, Poland, Denmark, Sweden, Svalbard, Ireland, Scotland, Wales, Iran, Kazakhstan and China. It is one of the last of the Agnostida order to survive.

<i>Ogygiocarella</i> Extinct genus of trilobites

Ogygiocarella Brongniart, 1822, is a genus of asaphid trilobites. It occurred during the Middle Ordovician.

Dolhir Formation

The Dolhir Formation is a geologic formation in Wales. It preserves fossils dating back to the Ordovician period.

<i>Bailiaspis</i> Genus of trilobites

Bailiaspis Resser, 1936, is a Middle Cambrian (Miaolingian) trilobite genus belonging to the Family Conocoryphidae Angelin, 1854. Within the Acado-Baltic region, the genus ranges from Wuliuan into Guzhangian age strata.

Palaeontology in Wales

Paleontology of Wales is paleontological research occurring in Wales.

References

  1. "†Gravicalymene Shirley 1936 (trilobite)". Paleobiology Database. Retrieved 21 August 2011.
  2. BRONGNIART, A. 1822. Les Trilobites. pp. 1-65, pls. 1-4 in: Histoire Naturelle des Crustacés Fossiles. Paris. F. G. Levrault, Libraire. 154 pp. 11 pls.
  3. SIVETER, D. J. 1977. The Middle Ordovician of the Oslo Region, Norway, 27. Trilobites of the Family Calymenidae. Norsk Geologisk Tidsskrift56: 335 - 396.
  4. SHIRLEY J. 1936. Some British trilobites of the family Calymenidae. Q. J. Geol. Soc. 92 (1–4): 384 – 422.
  5. PRICE, D. 1982. Calymene quadrata King, 1923 and allied species of trilobites from the Ashgill Series of North Wales. Geological Magazine, Volume 119, Issue 1 , January 1982 , pp. 57 - 66. DOI: https://doi.org/10.1017/S0016756800025656
  6. BANCROFT, B. B. 1949. Upper Ordovician trilobites of zonal value in south-east Shropshire (Edited by A. Lamont). Proc. R. Soc. (B), 136, pp. 291 - 315, pls. 9 - 11.
  7. LAMONT, A. 1946. Mr. B. B. Bancroft. Nature 157, p. 42. https://rdcu.be/cfPbu
  8. DEAN, W. T., 1962. The trilobite faunas of the Caradoc Series in the Cross Fell Inlier of Northern England. Bulletin British. Museum (natural History), Geology, 7 (3) pp. 65 - 134.
  9. DEAN, W. T., 1963, The Ordovician trilobite faunas of South Shropshire, III; British Mus. Nat. History Bull., Geology, v. 7, no. 8, pp. 215 - 254, pls. 37 - 46.
  10. CHATTERTON, B. D. E. 1971. Taxonomy and ontogeny of Siluro-Devonian trilobites from near Yass, New South Wales. Palaeontographica Abteilung A137 (1 - 3): 1 - 10.
  11. KOBAYASHI, T. & HAMADA, T., 1977. Devonian trilobites of Japan in comparison with Asian, Pacific and other faunas. Palaeontological Society of Japan, Special Papers, no. 20, pp. i–vii + 1–202, pls. 1–13. (Reference No. 0551)
  12. "Ohio University". Archived from the original on 2011-07-20. Retrieved 2011-01-06.
  13. Enrico de Lazaro: Fossils of New Trilobite Species Found in Tasmania, on:sci-news, Sep 28, 2020
  14. SMITH, P. M. & EBACH, M. C. 2020. A new Ordovician (Katian) calymenid, Gravicalymene bakeri sp. nov., from the Gordon Group, Tasmania, Australia. In:: Alcheringa: An Australasian Journal of Palaeontology, published online September 16, 2020; doi:10.1080/03115518.2020.1797874
  15. ROSS R. J. Jr, BARNES H. 1967. Some Middle Ordovician brachiopods and trilobites from the Basin Ranges, western United States. America: U.S. Govt. Print. Off.
  16. Hamed Ameri, Reza Arjmandzadeh & Khalil Ghoorcgi (2019): The peri-Gondwanian Early Silurian trilobites from Kopeh Dagh, Iran, Historical Biology. https://doi.org/10.1080/08912963.2019.1681420
  17. RUEDEMANN, R. 1926. The Utica and Lorraine Formations of New York. Pt. 2. N. Y. State Mus. Bull. No. 272.
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