The Cyatheaceae are a family of ferns, the scaly tree ferns, one of eight families in the order Cyatheales in the Pteridophyte Phylogeny Group classification of 2016. Alternatively, the family may defined much more broadly as the only family in the Cyatheales, with the PPG I family treated as the subfamily Cyatheoideae. The narrower circumscription is used in this article.
Alsophila aneitensis, synonym Cyathea aneitensis, is a species of tree fern native to Vanuatu and possibly New Caledonia. This species has an erect trunk up to 3 m tall. Fronds are bipinnate and may reach 2 m in length. The rachis and stipe are either very dark and smooth or have a few scales towards the base of the stipe. The scales are dark and narrow. Sori occur near the pinnule midvein and are covered by large, thin, fragile indusia. The closest relative of A. aneitensis appears to be Alsophila vieillardii. It can be distinguished from that species by its very dark stipes and frond bases.
Alsophila atropurpurea, synonym Cyathea atropurpurea, is a species of tree fern native to the islands of Luzon, Mindanao, Leyte and Mindanao in the Philippines, where it grows in forest at above 1000 m. The erect trunk is slender and may be up to 3 m tall. Fronds are bipinnate and 1–2 m long. Characteristically of this species, the final pair of pinnae are usually reduced and occur towards the base of the stipe. These, along with the stipe bases, are persistent and retained around the trunk long after withering. The stipe itself is dark and covered with scales, which are either small, dull and brown or large, dark and glossy. Sori occur near the midvein of fertile pinnules and lack indusia. Fertile pinnules are notably smaller than sterile ones.
Alsophila camerooniana, synonym Cyathea camerooniana, is a species of tree fern native to Sierra Leone, Cameroon, northern Angola and western Uganda, where it grows in montane forest at an altitude of 900–1200 m. The trunk is erect and 2–3 m tall. Fronds are pinnate and 2–3 m long. The rachis ranges in colour from dark to pale and has some hairs on the underside. The stipe is dark and is covered with scales throughout. Where scales have fallen, small warts are present. The scales are glossy brown and have a thin, dull edge. Sori occur at the forks of veins and are covered by thin indusia, which range in shape from cup- to saucer-like.
Alsophila coactilis, synonym Cyathea coactilis, is a rare species of tree fern known only from the southern highlands of Papua New Guinea, where it grows in alpine shrubland at an altitude of about 3000 m. The trunk is erect and 2–3 m tall. Fronds are bi- or tripinnate and usually 1–2 m long. Characteristically of this species, they occur in two whorls of about ten fronds each. The underside of the rachis is covered with small, pale scales. The stipe is covered with pale scales that have dark, narrow and fragile edges. Sori are round and are covered by thin indusia that are cup-like in appearance. They occur near the fertile pinnule midvein.
Alsophila cucullifera, synonym Cyathea cucullifera, is a species of tree fern native to eastern New Guinea, where it grows in montane forest at an altitude of about 2400 m. The trunk is erect and 2–3 m tall. Fronds are bi- or tripinnate and 2–3 m long. Characteristically of this species, they occur in two whorls of four to six fronds each. The stipe is warty and covered with scales. The scales are dark, glossy, have a narrow paler margin and are large towards the base. Sori occur near the fertile pinnule midvein and are covered by thin, pale brown indusia that are scale-like in appearance.
Alsophila eriophora, synonym Cyathea eriophora, is a species of tree fern native to eastern New Guinea, where it grows in wet ravine forest at an altitude of 1400–2000 m. The trunk is erect and 2–3 m tall. Fronds are bi- or tripinnate and 2–3 m long. The stipe is dark and covered with spines and scales. The scales are variable, being either small and pale or large with a dark apex. Sori occur near the fertile pinnule midvein and lack indusia.
Alsophila esmeraldensis is a species of tree fern which is endemic to Ecuador. It grows in coastal forest and forested slopes of the Andes.
Alsophila everta, synonym Cyathea everta, is a species of tree fern native to western New Guinea, where it grows in the edges of forest or in mossy forest at an altitude of 1400–2800 m. The trunk is erect and up to 5 m tall. Fronds are bi- or tripinnate and 1–1.5 m long. Numerous fronds are present at any one time and form a dense crown. The stipe has some spines towards the base and is covered in scales. The scales are pale or dark, glossy and have pale fringed edges. Sori occur near the fertile pinnule midvein. They are round and covered by thin, brown indusia that are cup-like in appearance.
Alsophila geluensis, synonym Cyathea geluensis, is a species of tree fern native to central and eastern New Guinea as well as the Louisiade Archipelago, where it grows in mossy forest. In New Guinea itself, plants grow at an altitude of 1000–2000 m, however they are present at lower elevations of 700–900 m on associated islands. The trunk of this tree fern is erect and may be 5 m tall or more. Fronds are bi- or tripinnate and 1–2.5 m long. They are usually about ten live fronds present in the crown at once. The stipe may be warty and/or have short spines as well as many scattered scales towards the base. These scales are pale to dark and have dull, fragile edges. Sori occur near the fertile pinnule midvein and are protected by pale, thin indusia. A. geluensis is a variable taxon and further study is needed to determine whether it does not in fact represent a species complex.
Gymnosphaera glabra, synonyms Alsophila glabra and Cyathea glabra, is a species of tree fern native to Borneo, western Java, Sumatra, the Lesser Sunda Islands, Sulawesi, and the Malay Peninsula, where it grows in lowland swamp forest and montane forest at an elevation of up to 1500 m. The trunk of this plant is erect and 2–4 m tall. Fronds are bi- or tripinnate and 1–2 m in length. Characteristically of this species, the lowest pinnae may be significantly reduced. The stipe is very dark and bears basal scales. These scales are dark, glossy and have a paler margin and fragile edges. Sori are produced in groups of one to three on fertile pinnule veins. They lack indusia.
Alsophila grevilleana, synonym Cyathea grevilleana, is a species of tree fern endemic to Jamaica, where it grows in moist gullies and on wooded hills in both calcareous and noncalcareous soils at an altitude of 200–1200 m. The trunk of this plant is erect, about 7 m tall, and 10–15 cm in diameter. It is characteristically clothed in old stipe bases, brown scales and blackish spines. Fronds are tripinnate, dark green in colour, and up to 4 m in length. The last pinnae are sometimes separated, forming a distinctive clump around the trunk apex. The rachis is yellow-brown and almost smooth. This species has a long, dark brown stipe with a few scattered spines. Sori are produced in four to six pairs along the pinnule midvein. They are protected by pale brown indusia that are cup-like in appearance.
Gymnosphaera denticulata, synonyms Alsophila acaulis, Alsophila denticulata and Cyathea hancockii, is a species of tree fern native to Japan, the Ryukyu Islands, Taiwan, southern China, Hong Kong, and Hainan. It grows in forest, on stream banks, and in forest margins at an elevation of about 600 m or higher. The specific epithet hancockii commemorates William Hancock (1847-1914), who collected numerous plants in Japan, China and Southeast Asia.
Alsophila × marcescens, synonym Cyathea × marcescens, commonly known as the skirted tree fern, is a tree fern endemic to the Cape Otway ranges in Victoria and Tasmania, Australia. It is a natural hybrid, apparently Alsophila australis × Alsophila cunninghamii. Large and Braggins (2004) note that it has characteristics midway between these two species. The spores of A. × marcescens are usually malformed although sterile. The trunk of this plant is erect and up to 10 m tall. Fronds may be bi- or tripinnate and 3–4 m in length. Dead fronds often persist, forming a characteristic skirt around the trunk. The stipe is thick, black and warty. The rachis and trunk are covered in shiny, dark brown scales. Sori are borne near the fertile pinnule midvein and are protected by thin indusia that are saucer-like in appearance.
Gymnosphaera hornei is a species of tree fern in the Cyatheaceae family.
Alsophila khasyana, synonym Cyathea khasyana, is a species of tree fern. Its natural distribution extends from India to Myanmar, although it is absent from Sri Lanka. A. khasyana grows in forest at an altitude of 1400–1700 m. The trunk of this plant is erect and 5–7 m tall. Fronds may be bi- or tripinnate and 2–3 m in length. A. khasyana has a long, dark stipe that is covered in numerous scales. These scales are dark and have broad, pale, fringed edges. Sori are borne near the midvein of fertile pinnules and lack indusia.
Alsophila brausei, synonym Cyathea hunsteiniana, is a species of tree fern endemic to eastern New Guinea, where it grows in rain forest at an altitude of 1300–2000 m. This species has a slender, erect trunk up to 1 m tall and about 3 cm in diameter. Fronds are bipinnate and approximately 1 m in length. Scattered scales cover the stipe. They are dark brown in colouration and have a broad paler margin and fragile edges. Round sori are borne one or two per fertile pinnule segment. The sori are protected by deep, firm indusia that are cup-like in appearance.
Alsophila imbricata, synonym Cyathea imbricata, is a species of tree fern endemic to Western New Guinea, where it grows in open forest at an altitude of 3240 m. The trunk of this plant is erect and approximately 2 m tall. Fronds may be bi- or tripinnate and are usually less than 1 m in length. The stipe is dark, spiny, and covered with caducous scales. These scales are glossy brown in colouration and have a paler margin and fragile edges. Sori are borne in groups of one to four per pinnule lobe. They are protected by firm indusia.
Alsophila javanica, synonym Cyathea javanica, is a species of tree fern native to western Java and Sumatra, in Indonesia. It grows in rain forest and on riverbanks at an altitude of 250–1500 m.
Alsophila junghuhniana, synonym Cyathea junghuhniana, is a species of tree fern native to southern and central Sumatra and western Java, where it grows in forest at an altitude of 1000–2000 m. The trunk of this species is erect and may be 2 m tall or more. Fronds are tripinnate and up to about 3 m long. The stipes of this species are persistent and form a skirt around the trunk. They are brown, spiny, and covered with glossy, dark brown scales. Sori are borne near the midvein of fertile pinnules and are protected by thin indusia.
