| Hantkenina Temporal range: | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Clade: | Diaphoretickes |
| Clade: | SAR |
| Phylum: | Retaria |
| Infraphylum: | Foraminifera |
| Class: | Globothalamea |
| Order: | Rotaliida |
| Family: | † Hantkeninidae |
| Genus: | † Hantkenina Cushman, 1924 |
| Species | |
| |
Hantkenina is a genus of planktonic foraminifera that lived from the Middle Eocene up to late Eocene, cirka 49 Ma-33.9 Ma. There have been 11 morphospecies described, including one of Cribrohantkenina [1]
Hantkenina is a genus of foraminifera. Hantkeninids are well-known biostratigraphic index fossils. [2] They produce a test ("shell") of calcite (CaCO3). Hantkeninids evolved from Clavigerinella caucasica . [1] Hantkenina is highly distinctive from other planktonic foraminifers as they are characterized by planispiral coiling and their hollow, slender extensions of each chamber known as tubulospines. [2]
The structure of this unique feature vary between individuals, and between species of hantkenina. All hantkeninas species retained tubulospines through their evolution. This indicates that it served a useful purpose for this particular form of foraminifera. The function of the tubulospines are unknown, however, reproduction and feeding purposes have been suggested as possible functions. [3]
The first hantkenids (Hantkenina mexicana) lived in deep planktonic environments with minimum oxygen levels. However, isotopes suggests they migrated to fully oxygenated shallower waters at about 43.8 Ma. [2] During the Eocene the climate went through climate changes. The peak of the Cenozoic warmth occurred during the Early Eocene. Afterwards, global cooling began, persisting to Early Oligocene. [3] The first large ice caps appeared in Antarctica during this time. [4] This climate change also affected deep-water environments.
The cooling process caused gradual changes in the structure of the water column. As the water cooled, the rates of remineralization of organic matter at the near surface water altered. A consequence of this global cooling was that bacterial metabolic rates at the upper water column slowed down and allowed sinking organic matter to descend further to deeper environments in the ocean. This made the oxygen minimum zone, less intense and more spread out. New niches for deep-dwelling zooplankton that were able to tolerate low levels of oxygen opened up. Pearson and Coxall (2013) speculate that the evolution of Clavigerinella and hantkenina was related to this global cooling, [3] as well as pulses of deep-water anoxia [5]
The Cenozoic is Earth's current geological era, representing the last 66 million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configuration of continents. It is the latest of three geological eras since complex life evolved, preceded by the Mesozoic and Paleozoic. It started with the Cretaceous–Paleogene extinction event, when many species, including the non-avian dinosaurs, became extinct in an event attributed by most experts to the impact of a large asteroid or other celestial body, the Chicxulub impactor.
The Eocene Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name Eocene comes from the Ancient Greek ἠώς and καινός and refers to the "dawn" of modern ('new') fauna that appeared during the epoch.
The Miocene is the first geological epoch of the Neogene Period and extends from about 23.03 to 5.333 million years ago (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words μείων and καινός and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene.
The Oligocene is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present. As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from the Ancient Greek ὀλίγος and καινός, and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period.
The Paleogene is a geologic period and system that spans 43 million years from the end of the Cretaceous Period 66 million years ago (Mya) to the beginning of the Neogene Period 23.03 Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon. The earlier term Tertiary Period was used to define the span of time now covered by the Paleogene Period and subsequent Neogene Period; despite no longer being recognized as a formal stratigraphic term, "Tertiary" still sometimes remains in informal use. Paleogene is often abbreviated "Pg".
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Foraminifera are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an external shell of diverse forms and materials. Tests of chitin are believed to be the most primitive type. Most foraminifera are marine, the majority of which live on or within the seafloor sediment, while a smaller number float in the water column at various depths, which belong to the suborder Globigerinina. Fewer are known from freshwater or brackish conditions, and some very few (nonaquatic) soil species have been identified through molecular analysis of small subunit ribosomal DNA.
The Deep Sea Drilling Project (DSDP) was an ocean drilling project operated from 1968 to 1983. The program was a success, as evidenced by the data and publications that have resulted from it. The data are now hosted by Texas A&M University, although the program was coordinated by the Scripps Institution of Oceanography at the University of California, San Diego. DSDP provided crucial data to support the seafloor spreading hypothesis and helped to prove the theory of plate tectonics. DSDP was the first of three international scientific ocean drilling programs that have operated over more than 40 years. It was followed by the Ocean Drilling Program (ODP) in 1985, the Integrated Ocean Drilling Program in 2004 and the present International Ocean Discovery Program in 2013.
Oceanic anoxic events or anoxic events (anoxia conditions) describe periods wherein large expanses of Earth's oceans were depleted of dissolved oxygen (O2), creating toxic, euxinic (anoxic and sulfidic) waters. Although anoxic events have not happened for millions of years, the geologic record shows that they happened many times in the past. Anoxic events coincided with several mass extinctions and may have contributed to them. These mass extinctions include some that geobiologists use as time markers in biostratigraphic dating. On the other hand, there are widespread, various black-shale beds from the mid-Cretaceous which indicate anoxic events but are not associated with mass extinctions. Many geologists believe oceanic anoxic events are strongly linked to the slowing of ocean circulation, climatic warming, and elevated levels of greenhouse gases. Researchers have proposed enhanced volcanism (the release of CO2) as the "central external trigger for euxinia."
TEX86 is an organic paleothermometer based upon the membrane lipids of mesophilic marine Nitrososphaerota (formerly Marine Group 1 Crenarchaeota).
Paleoceanography is the study of the history of the oceans in the geologic past with regard to circulation, chemistry, biology, geology and patterns of sedimentation and biological productivity. Paleoceanographic studies using environment models and different proxies enable the scientific community to assess the role of the oceanic processes in the global climate by the re-construction of past climate at various intervals. Paleoceanographic research is also intimately tied to paleoclimatology.
The Eocene–Oligocene extinction event, also called the Eocene-Oligocene transition or Grande Coupure, is the transition between the end of the Eocene and the beginning of the Oligocene, an extinction event and faunal turnover occurring between 33.9 and 33.4 million years ago marked by large-scale extinction and floral and faunal turnover. Most of the affected organisms were marine or aquatic in nature. They included the last of the ancient ungulates, the "condylarths".
The Paleocene, or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek παλαιός palaiós meaning "old" and the Eocene Epoch, translating to "the old part of the Eocene".
The Cretaceous Thermal Maximum (CTM), also known as Cretaceous Thermal Optimum, was a period of climatic warming that reached its peak approximately 90 million years ago (90 Ma) during the Turonian age of the Late Cretaceous epoch. The CTM is notable for its dramatic increase in global temperatures characterized by high carbon dioxide levels.
The Cenomanian-Turonian boundary event, also known as the Cenomanian-Turonian extinction, Cenomanian-Turonian oceanic anoxic event, and referred to also as the Bonarelli event, was one of two anoxic extinction events in the Cretaceous period. The Cenomanian-Turonian oceanic anoxic event is considered to be the most recent truly global oceanic anoxic event in Earth's geologic history. Selby et al. in 2009 concluded the OAE 2 occurred approximately 91.5 ± 8.6 Ma, though estimates published by Leckie et al. (2002) are given as 93–94 Ma. The Cenomanian-Turonian boundary has been refined in 2012 to 93.9 ± 0.15 Ma. There was a large carbon cycle disturbance during this time period, signified by a large positive carbon isotope excursion. However, apart from the carbon cycle disturbance, there were also large disturbances in the nitrogen, oxygen, and sulphur cycles of the ocean.
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