Inchoatia haussknechti

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Inchoatia haussknechti
Inchoatia haussknechti shell.png
two views of the shell of Inchoatia haussknechti haussknechti from its type description by Oskar Boettger
Scientific classification
Kingdom:
Animalia
Phylum:
Mollusca
Class:
Gastropoda
(unranked):
clade Heterobranchia

clade Euthyneura
clade Panpulmonata
clade Eupulmonata
clade Stylommatophora

informal group

Contents

Sigmurethra
Superfamily:
Clausilioidea
Family:
Clausiliidae
Subfamily:
Alopiinae
Tribe:
Medorini
Genus:
Inchoatia
Species:
I. haussknechti
Binomial name
Inchoatia haussknechti
(O. Boettger, 1886) [1]
Synonyms

Clausilia (Albinaria) Haussknechti Boettger, 1886

Inchoatia haussknechti is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Clausiliidae, the door snails.

Description

The shell is dark horny brownish, very finely striated, slightly coarser at the cervix. [2] The shell has 9-11 whorls. [2] The cervix is with strong basal and dorsal keels and a basal forrow in between, also inside the aperture. [2] The columellaris is not very prominent, the frontal upper palatalis is short, basalis is short, subcolumellaris is visible in an oblique view. [2]

The width of the shell is 3-3.5 mm. [2] The height of the shell is 13–17 mm. [2]

Distribution

Inchoatia haussknechti is distributed in limestone areas from south Epirus to Central Greece. [2] [3]

Distribution of Inchoatia haussknechti in Greece: yellow dot = Inchoatia haussknechti haussknechti red dot = Inchoatia haussknechti orina violet dot = Inchoatia haussknechti alticola dark blue dot = Inchoatia haussknechti hiltrudae light blue dot = Inchoatia haussknechti refuga Inchoatia haussknechti map.png
Distribution of Inchoatia haussknechti in Greece:
yellow dot = Inchoatia haussknechti haussknechti
red dot = Inchoatia haussknechti orina
violet dot = Inchoatia haussknechti alticola
dark blue dot = Inchoatia haussknechti hiltrudae
light blue dot = Inchoatia haussknechti refuga

Subspecies

According to the Gittenberger & Uit de Weerd (2009) [3] the species Inchoatia haussknechti include 6 subspecies:

Inchoatia haussknechti haussknechti

Synonyms include: [3]

Diagnosis: The entire teleoconch is with sharp, mostly whitened riblets. The parietal side of the apertural border is not protruding. Lamella columellaris is shortly protruding into the aperture. Lamella parietalis is moderately long, i.e. reaching slightly further than the lamella spiralis. The lunella is prominent. [3]

Range: The type locality is Greece, Thessalia, Karditsa: northern side of the Voutsikaki Mts (= 25 km WSW of Karditsa). [1] [3] The type locality ‘Gion Skala’, apparently a high, narrow passage, cut through the rocks at the northern side of the Voutsikaki Mountains, could be located on the basis of the detailed excursion report by Stussiner, in Boettger (1886: 47). [1] [3] The disjunct range of this subspecies is built up by three populations in the eastern Pindos Mountains. [3]

Notes: The population that was discovered recently at an isolated limestone outcrop near Mesochori  [ el ], is located 45 km SE of the type locality. [3] The Mount Oeta, where according to Nordsieck (1972, [4] 1974 [5] ) the same subspecies occurs (see Inchoatia haussknechti orina), is situated 35 km SE of Mesochori and 80 km SE of the Voutsikaki Mts. It should be investigated whether these disjunctions, which are quite extreme when compared with the distributional patterns of the other subspecies, are not correlated with any taxonomically relevant differentiation. [3]

Inchoatia haussknechti alticola

Synonyms include: [3]

Diagnosis: Initial teleoconch whorls with lengthened white papillae, without regular riblets; parietal side of the apertural border not protruding; lamella parietalis short; lamella columellaris (very) low; lunella rather prominent. [3]

Range: This subspecies is known from the Mt. Timfristos (= Veluchi) area, north of Karpenisi. [3] The type locality is 4 km north of Karpenisi, Mt. Timfristos ski resort, 1900 m alt. [3]

Initially (Nordsieck, 1972) [4] this form was not separated from Inchoatia haussknechti semilaevis. [3]

Inchoatia haussknechti hiltrudae

Synonyms include: [3]

Diagnosis: Initial teleoconch whorls with often whitened riblets, which become more irregular and are often not whitened on the lower whorls; parietal side of the apertural border usually protruding; lamella columellaris clearly protruding into the aperture; lamella parietalis moderately long, i.e. reaching slightly further than the lamella spiralis; lunella prominent. [3]

Range: This subspecies is only known from the type locality: Greece, Thessalia, Trikala: 1 km W of Pyli (= Pili), near the classical bridge, 300 m alt. [3]

Inchoatia haussknechti orina

Synonyms include: [3]

Material.― Greece, Fthiotida/Fokida, Mt. Oiti (= Iti), c. FH1097 (but see ‘Range’).

Diagnosis.― See the notes.

Range: The type locality ‘Koraki Besa im Oetagebirge’ (Westerlund, 1894: 175) could not be located. The Mt Oeta is nowadays called Oiti or Iti Oros (situated 20 km SW of Lamia). Nordsieck (1974: 148) [5] suggested that ‘Koraki Besa’ refers to the Korax Mtn (= Korakas) in the Vardousia Mts (about 35 km SW of Lamia), c. 15 km SW of the Oiti Oros. The area should be visited to get more certainty. [3] The material examined by Gittenberger & Uit de Weerd (2009) is listed as: "Greece, Fthiotida/Fokida, Mt. Oiti (= Iti), c. FH1097". [3]

Neither a detailed description nor a figure is available, but a lectotype has been selected by Nordsieck (1972: 16). [3] [4] Nordsieck (1972: 16; 1974: 148) [4] [5] considered this taxon a synonym of the nominate subspecies, whereas Zilch (1981: 125) [7] and Nordsieck (2007: 48) [8] listed it as a subspecies, without adding more data. [3]

Inchoatia haussknechti refuga

Synonyms include: [3]

Diagnosis: The entire teleoconch is with sharp, mostly whitened riblets. The parietal side of the apertural border is protruding. Lamella columellaris is shortly protruding into the aperture. The lower part of the lunella is obsolete. [3] The subspecies was characterized, with the designation of a lectotype, by Nordsieck (1972). [3] [4]

Range: Only known from the type locality. ‘Katafiyi (800 m)’ and ‘Katafiyi (2000 m)’, mentioned by Nordsieck (1972), [4] refer to Katafigio (= Katafiyion) at the western foot of the Tschumerka Mts. [3]

Inchoatia haussknechti semilaevis

Synonyms include: [3]

Diagnosis: Only the initial whorls of the teleoconch and the cervical part with sharp, partly whitened riblets; parietal side of the apertural border not protruding; lamella columellaris shortly protruding into the aperture; lunella prominent. [3]

Range: The type locality of this subspecies is indicated in a somewhat misleading way because the Mt Veluchi (= Velouhi) or Mt Timfristos is situated a few km N of Karpenisi, where Inchoatia haussknechti alticola occurs, whereas “Kaljakuda” refers to Mt Kaliakouda, 16 km SSW of Karpenisi. The localities ‘Karpenission’ and ‘Timfristos-Geb. 13 km n. Karpenission’, cited for this subspecies by Nordsieck (1972), [4] should be transferred to Inchoatia haussknechti alticola, though only the latter record is repeated (in a slightly different wording) by Nordsieck (1974). [3] [5]

Related Research Articles

<span class="mw-page-title-main">Clausiliidae</span> Family of gastropods

Clausiliidae, also known by the common name door snails, is a taxonomic family of small, very elongate, mostly left-handed, air-breathing land snails, sinistral terrestrial pulmonate gastropod mollusks.

This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary of leaf morphology. For other related terms, see Glossary of phytopathology, Glossary of lichen terms, and List of Latin and Greek words commonly used in systematic names.

<span class="mw-page-title-main">Cro-Magnon</span> Earliest anatomically modern humans in Europe

Cro-Magnons or European early modern humans (EEMH) were the first early modern humans to settle in Europe, migrating from western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They interacted and interbred with the indigenous Neanderthals of Europe and Western Asia, who went extinct 40,000 to 35,000 years ago. The first wave of modern humans in Europe left no genetic legacy to modern Europeans; however, from 37,000 years ago a second wave succeeded in forming a single founder population, from which all subsequent Cro-Magnons descended and which contributes ancestry to present-day Europeans. Cro-Magnons produced Upper Palaeolithic cultures, the first major one being the Aurignacian, which was succeeded by the Gravettian by 30,000 years ago. The Gravettian split into the Epi-Gravettian in the east and Solutrean in the west, due to major climatic degradation during the Last Glacial Maximum (LGM), peaking 21,000 years ago. As Europe warmed, the Solutrean evolved into the Magdalenian by 20,000 years ago, and these peoples recolonised Europe. The Magdalenian and Epi-Gravettian gave way to Mesolithic cultures as big game animals were dying out and the Last Glacial Period drew to a close.

<i>Inchoatia</i> Genus of gastropods

Inchoatia is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Clausiliidae, the door snails.

<i>Inchoatia megdova</i> Species of gastropod

Inchoatia megdova is a species of air-breathing land snail, terrestrial pulmonate gastropod mollusk in the family Clausiliidae, the door snails.

<i>Inchoatia inchoata</i> Species of gastropod

Inchoatia inchoata is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Clausiliidae, the door snails.

<i>Atractophaedusa smithi</i> Species of gastropod

Atractophaedusa smithi is a species of air-breathing land snail, a terrestrial gastropod mollusk in the family Clausiliidae, the door snails.

References

This article incorporates CC-BY-3.0 text from reference [3] and public domain text from the reference. [2]

  1. 1 2 3 PD-icon.svg (in German) Boettger O. (1886). "Aufzählung der in Thessalien gesammelten Schnecken und Muscheln II". Jahrbücher der Deutschen Malakozoologischen Gesellschaft13: 50-73. description is on the page 61, Table 2, figure 6a-c. Frankfurt am Main.
  2. 1 2 3 4 5 6 7 8 "Species summary for Carinigera haussknechti". AnimalBase, last modified 14 February 2009, accessed 17 August 2010.
  3. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 Gittenberger E. & Uit de Weerd D. R. (2009). "Summarizing data on the Inchoatia taxa, including Inchoatia megdova bruggeni subspec. nov. (Gastropoda, Pulmonata, Clausiliidae)". Zoologische Mededelingen 83 http://www.zoologischemededelingen.nl/83/nr03/a08 Archived 2014-01-16 at the Wayback Machine
  4. 1 2 3 4 5 6 7 8 9 Nordsieck H. (1972). "Zur Anatomie und Systematik der Clausilien, XI. Neue Formen und taxonomische Revision einiger Gruppen der Alopiinae". Archiv für Molluskenkunde102: 1-51.
  5. 1 2 3 4 5 6 Nordsieck H. (1974). "Zur Anatomie und Systematik der Clausilien, XV. Neue Clausilien der Balkan-Halbinsel (mit taxonomischer Revision einiger Gruppen der Alopiinae und Baleinae)". Archiv für Molluskenkunde104: 123-170.
  6. 1 2 Westerlund C. A. (1894). "Specilegium malacologicum". Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft26: 163-177.
  7. 1 2 Zilch A. (1981). "Die Typen und Typoide des Natur-Museums Senckenberg, 65: Mollusca: Clausiliidae (6): Alopiinae (4): Montenegrinini". Archiv für Molluskenkunde111: 123-145.
  8. ↑ Nordsieck H. (2007). Worldwide door snails (Clausiliidae), recent and fossil. 1-214. ConchBooks, Hackenheim.
  9. ↑ Boettger O. (1889). "Ein Paar neue Fundorte griechischer Landschnecken". Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft21: 23-26.
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