Kokartus Temporal range: Middle Jurassic, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Amphibia |
Family: | † Karauridae |
Genus: | † Kokartus Nessov, 1988 |
Species: | †K. honorarius |
Binomial name | |
†Kokartus honorarius Nessov, 1988 | |
Kokartus is an extinct genus of prehistoric stem-group salamander (Caudata) from the Middle Jurassic Balabansai Formation of Kyrgyzstan. [1]
The absence of clear lines of arrested growth and annuli in long bones suggests that the animals lived in an environment with stable local conditions. [2] Like other members of Karauridae, it is thought to have been neotenic. [3] Kokartus is thought to have fed using suction feeding via the enlargement of the buccal cavity on small fish and invertebrates, with the well developed palatal dentition (teeth on the roof of the mouth) and marginal teeth helping to grasp prey. [4]
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela from the group Caudata. Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm.
The Lissamphibia is a group of tetrapods that includes all modern amphibians. Lissamphibians consist of three living groups: the Salientia, the Caudata, and the Gymnophiona.
The Cryptobranchidae are a family of fully aquatic salamanders commonly known as the giant salamanders. They include some of the largest living amphibians. The family is native to China, Japan, and the eastern United States. They constitute one of two living families—the other being the Asiatic salamanders belonging to the family Hynobiidae—within the Cryptobranchoidea, one of two main divisions of living salamanders.
The Caudata are a group of amphibians containing the extant salamanders and all extinct species of amphibians more closely related to salamanders than to frogs. They are typically characterized by a superficially lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults.
Caecilians are a group of limbless, vermiform (worm-shaped) or serpentine (snake-shaped) amphibians with small or sometimes nonexistent eyes. They mostly live hidden in soil or in streambeds, and this cryptic lifestyle renders caecilians among the least familiar amphibians. Modern caecilians live in the tropics of South and Central America, Africa, and southern Asia. Caecilians feed on small subterranean creatures such as earthworms. The body is cylindrical and often darkly coloured, and the skull is bullet-shaped and strongly built. Caecilian heads have several unique adaptations, including fused cranial and jaw bones, a two-part system of jaw muscles, and a chemosensory tentacle in front of the eye. The skin is slimy and bears ringlike markings or grooves and may contain scales.
Karaurus is an extinct genus of stem-group salamander (Caudata) from the Middle to Late Jurassic (Callovian–Kimmeridgian) Karabastau Formation of Kazakhstan. It is one of the oldest salamanders known.
The Salamandroidea are a suborder of salamanders, referred to as advanced salamanders. The members of the suborder are found worldwide except for Antarctica, sub-Saharan Africa, and Oceania. They differ from suborder Cryptobranchoidea as the angular and prearticular bones in their lower jaws are fused, their trunk ribs are bicapitate, and all members use internal fertilization. The female is fertilized by means of a spermatophore, a sperm-containing cap placed by the male in her cloaca. The sperm is stored in spermathecae on the roof of the cloaca until it is needed at the time of oviposition.
The Albanerpetontidae are an extinct family of small amphibians, native to the Northern Hemisphere during the Mesozoic and Cenozoic. The only members of the order Allocaudata, they are thought to be allied with living amphibians belonging to Lissamphibia. Despite a superficially salamander-like bodyform, their anatomy is strongly divergent from modern amphibians in numerous aspects. The fossil record of albanerpetontids spans over 160 million years from the Middle Jurassic to the beginning of the Pleistocene, about 2.13–2 million years ago.
Chunerpeton tianyiensis is an extinct species of salamander from the Late Jurassic Daohugou Beds in Ningcheng County, Nei Mongol, China. It is the only species classified under the genus Chunerpeton, which means "early creeping animal". It was a small animal measuring 18 cm in length. It was neotenic, with the retention of external gills into adulthood. In the original description it was placed in Cryptobranchidae, which contains modern giant salamanders. A redescription published in 2020 found it to be a stem-group caudatan outside the crown group of modern salamanders. A 2021 study found it to be a member of Cryptobranchoidea outside of Cryptobranchidae. In 2022 a more extensive analysis, with greater character and taxon sampling, recovered Chunerpeton tianyiense as a stem-group caudatan, outside the crown group of modern salamanders, and associated with Beiyanerpeton jianpingensis and Qinglongtriton gangouensis.
Eoscapherpeton is an extinct genus of giant salamander, known from the Late Cretaceous of Central Asia. Fossils have been found in the Cenomanian aged Khodzhakul Formation and Dzharakuduk Formation, Turonian aged Bissekty Formation and the Coniacian-Santonian aged Aitym Formation of Uzbekistan, the Santonian aged Yalovach Formation of Tajikistan, and the Santonian-lower Campanian aged Bostobe Formation and Campanian aged Darbasa Formation of Kazakhstan.
Iridotriton is an extinct genus of prehistoric salamander known from a fossil found in stratigraphic zone 6 of the late Jurassic Morrison Formation in the Dinosaur National Monument. One species has been described, Iridotriton hechti. It is likely a member of Cryptobranchoidea.
Hylaeobatrachus is an extinct genus of prehistoric salamander, known from the Early Cretaceous of Europe. The type species H. croyii is known from the Sainte-Barbe Clays Formation at the Iguanodon locality of Belgium, and was described by Louis Dollo. An unnamed Hylaeobatrachus-like taxon has also been reported from Las Hoyas, Spain. Both localities are of Barremian age. Hylaeobatrachus belongs to the crown group of modern salamanders, though its exact relationship with modern salamander groups is uncertain. It was neotenic, llike some modern salamanders.
Marmorerpeton is an extinct genus of prehistoric stem group-salamanders that lived in Britain during the Bathonian stage of the Middle Jurassic. They are among the oldest known salamanders. Two species were named when the genus was first described by Susan E. Evans et al. in 1988, M. freemani, and M. kermacki, from fragmentary remains found via screenwashing in the Forest Marble Formation of England. Due to the size of their osteocytic lacunae suggesting a large genome size and some morphological characters, like the presence of calcified cartilage in the medulla of its humerus, it was assumed that Marmorerpeton was neotenic. New more complete remains of a new species M. wakei were described in 2022 from the Kilmaluag Formation of the Isle of Skye, Scotland. These conclusively demonstrated that Marmorerpeton was neotenic, and was a member of the family Karauridae, with the other two members of the family, Karaurus and Kokartus being known from the Middle-Late Jurassic of Central Asia. The teeth appear to have been weakly pedicellate.
The Balabansai Formation is a geological formation in Kazakhstan, Kyrgyzstan and Uzbekistan whose strata date back to the Bathonian and Callovian stages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from the formation. The lithology primarily consists of variegated sandstones, siltstones, claystones, and rare gravels and marls. Many taxa have been found in the formation, including amphibians and mammals
The Itat Formation is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from the formation, including the proceratosaurid Kileskus, as well as fish, amphibians, mammals and many other vertebrates. The formation is noted for bearing significant coal reserves, with large open pit coal mines extracting lignite from the unit currently in operation.
The Karauridae are a family of stem-group salamanders (Caudata) that are known from the Middle Jurassic to Early Cretaceous in Central Asia, Northern Asia and Western Europe. The family includes four members: Karaurus from the Middle-Late Jurassic Karabastau Formation of Kazakhstan, Kokartus from the Middle Jurassic Balabansai Formation of Kyrgyzstan, Marmorerpeton from the Middle Jurassic Forest Marble Formation of England and Kilmaluag Formation of Scotland, and Kuzbassia from the Early Cretaceous (Aptian) Ilek Formation. The members are some of the oldest known salamanders. The family is united by several morphological characters, including sculptured skull roof bones. Like some modern salamanders, karaurids were neotenic. Members of the family likely fed via suction feeding on small fish and invertebrates. The Early Cretaceous Siberian Kulgeriherpeton has been suggested to be a karaurid by some authors.
The Batylykh Formation is a geological formation in Yakutia, Russia. It is of an uncertain Early Cretaceous age, probably dating between the Berriasian and the Barremian. It is the oldest unit of the 1,500 metres (4,900 ft) thick Sangar Series within the Vilyuy syneclise. The mudstones, sandstones and shales of the formation were deposited in a fluvial to lacustrine environment.
Triassurus is an extinct genus of amphibian, and the oldest member of Caudata. It is known from the Middle to Upper Triassic (Ladinian-Carnian) aged Madygen Formation in Kyrgyzstan. The type species is T. sixtelae.
This list of fossil amphibians described in 2018 is a list of new taxa of fossil amphibians that were described during the year 2018, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2018.
This list of fossil amphibians described in 2020 is a list of new taxa of fossil amphibians that were described during the year 2020, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2020.