Ciconiidae is a family of heavy-bodied, large-billed wading birds in the monotypic order Ciconiiformes. Most species in the family are called storks, although some have different common names: two species in the genus Anastomus are known as openbills, two from the genus Leptoptilos are called adjutants, and three species are called jabiru. [1] Storks are found in tropical and subtropical habitats around the world, mostly inhabiting wetlands and marshes, although some also inhabit forests and savannah. [2] [3] They are large birds with long legs, stout bills, and variable featherless patches on the head. The beaks of storks can be highly specialised depending on their function, as exemplified by the openbills, whose bills possess a gap between the mandibles to help capture freshwater snails. [4]
Storks are threatened by habitat loss, pesticide use, and intentional hunting. [2] Four species of stork—the milky stork, oriental stork, greater adjutant, and Storm's stork—are classified as being endangered on the IUCN Red List, while a further three are considered near-threatened and one is considered vulnerable. However, the conservation statuses of many species of stork seem to be based on mistaken assumptions and limited data, such as underestimating the ability of species to use agricultural land. Many species may consequently warrant a data deficient listing until more information about their ecology is available. [5]
There are currently 20 extant species of stork recognised by the International Ornithologists' Union, distributed among 6 genera, 1 of which is monotypic. [1] The order Ciconiiformes previously also included the herons (Ardeidae) and ibises and spoonbills (Threskiornithidae), but these have since been reassigned to other orders. [2] The storks are most diverse in Afro-Eurasia, especially in Africa and Asia, and only three species, including the wood stork, are known from the Americas. [3] [5] Many species of fossil stork are also known from the Oligocene onwards; however, their exact number and taxonomy are unsettled due to ongoing discoveries. [6]
Conservation status | |
---|---|
EX | Extinct (0 species) |
EW | Extinct in the wild (0 species) |
CR | Critically Endangered (0 species) |
EN | Endangered (4 species) |
VU | Vulnerable (1 species) |
NT | Near threatened (3 species) |
LC | Least concern (12 species) |
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the stork's range is provided. Ranges are based on the IOC World Bird List for that species unless otherwise noted. Population estimates are of the number of mature individuals and are taken from the IUCN Red List.
This list follows the taxonomic treatment (designation and order of species) and nomenclature (scientific and common names) of version 13.2 of the IOC World Bird List. [1] Where the taxonomy proposed by the IOC World Bird List conflicts with the taxonomy followed by the IUCN [lower-alpha 1] or the 2023 edition of The Clements Checklist of Birds of the World, [8] the disagreement is noted next to the species's common name (for nomenclatural disagreements) or scientific name (for taxonomic disagreements).
The International Ornithologists' Union (IOU) recognises 20 species of storks in six genera. [1] This list does not include hybrid species, extinct prehistoric species, or putative species not yet accepted by the IOU. Ciconiidae has traditionally been divided into three tribes: Mycteriini, containing Mycteria and Anastomus , Ciconiini, containing Ciconia , and Leptoptilini, containing Leptoptilos , Ephippiorhynchus, and Jabiru . However, the exact division of these genera among these tribes is contentious and different authors recognise from two to four tribes. [3] [4] Recent genetic studies incorporated by the IOU have suggested that Mycteriini and Leptoptilini are paraphyletic (not including all of a common ancestor's descendants), and that Ephippiorhynchus and Jabiru should be moved into their own tribe, Ephippiorhynchini. [3]
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Phylogeny recognised by the International Ornithologists' Union, based partially on de Sousa and colleagues, 2023. [1] [3] |
Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
---|---|---|---|
African openbill | A. lamelligerus Temminck, 1823 Two subspecies
| Africa | LC
|
Asian openbill | A. oscitans (Boddaert, 1783) | India and Southeast Asia | LC
|
Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
---|---|---|---|
Marabou stork [lower-alpha 2] | L. crumenifer (Lesson, R. P., 1831) | Sub-Saharan Africa | LC
|
Lesser adjutant | L. javanicus (Horsfield, 1821) | South and Southeast Asia | VU
|
Greater adjutant | L. dubius (Gmelin, J. F., 1789) | Northern India | EN
|
Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
---|---|---|---|
Wood stork | M. americana Linnaeus, 1758 | Americas, from southeastern United States south to northern Argentina | LC
|
Yellow-billed stork | M. ibis (Linnaeus, 1766) | Sub-Saharan Africa | LC
|
Painted stork | M. leucocephala (Pennant, 1769) | India and Southeast Asia | NT
|
Milky stork | M. cinerea (Raffles, 1722) | Southeast Asia | EN
|
Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
---|---|---|---|
Jabiru | J. mycteria (Lichtenstein, M. H. C., 1819) | Central and South America | LC
|
Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
---|---|---|---|
Saddle-billed stork [lower-alpha 3] | E. senegalensis (Shaw, 1800) | Sub-Saharan Africa | LC
|
Black-necked stork | E. asiaticus (Latham, 1790) Two subspecies
| India, Southeast Asia, New Guinea, and Australia | NT
|
Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
---|---|---|---|
Abdim's stork | C. abdimii Lichtenstein, M. H. C., 1823 | Africa | LC
|
Asian woolly-necked stork [lower-alpha 4] | C. episcopus (Boddaert, 1783) Two subspecies
| South and Southeast Asia | NT
|
Storm's stork | C. stormi (Blasius, W., 1896) | Malay Peninsula, Borneo, Sumatra, and nearby islands | EN
|
Black stork | C. nigra (Linnaeus, 1758) | Widespread through Africa, Asia, and Europe | LC
|
Maguari stork | C. maguari (Gmelin, J. F., 1789) | South America | LC
|
African woolly-necked stork [lower-alpha 5] | C. microscelis Gray, G. R., 1848 | Sub-Saharan Africa | LC
|
White stork | C. ciconia (Linnaeus, 1758) Two subspecies
| Widely throughout Africa and Paleartic of Asia and Europe | LC
|
Oriental stork | C. boyciana Swinhoe, 1873 | Siberia, China, Korea, and Japan | EN
|
Storks are large, long-legged, long-necked wading birds with long, stout bills. They belong to the family Ciconiidae, and make up the order Ciconiiformes. Ciconiiformes previously included a number of other families, such as herons and ibises, but those families have been moved to other orders.
The marabou stork is a large wading bird in the stork family Ciconiidae native to sub-Saharan Africa. It breeds in both wet and arid habitats, often near human habitation, especially landfill sites. It is sometimes called the "undertaker bird" due to its shape from behind: cloak-like wings and back, skinny white legs, and sometimes a large white mass of "hair". It has the largest wingspan of any land bird, with an average of 2.6 metres (8.5 ft) and some recorded examples of up to 3.2 metres (10 ft).
The lesser adjutant is a large wading bird in the stork family Ciconiidae. Like other members of its genus, it has a bare neck and head. It is however more closely associated with wetland habitats where it is solitary and is less likely to scavenge than the related greater adjutant. It is a widespread species found from India through Southeast Asia to Java.
The black-necked stork is a tall long-necked wading bird in the stork family. It is a resident species across the Indian Subcontinent and Southeast Asia with a disjunct population in Australia. It lives in wetland habitats and near fields of certain crops such as rice and wheat where it forages for a wide range of animal prey. Adult birds of both sexes have a heavy bill and are patterned in white and irridescent blacks, but the sexes differ in the colour of the iris with females sporting yellow irises and males having dark-coloured irises. In Australia, it is known as a jabiru although that name refers to a stork species found in the Americas. It is one of the few storks that are strongly territorial when feeding and breeding.
The jabiru is a large stork found in the Americas from Mexico to Argentina, except west of the Andes. It sometimes wanders into the United States, usually in Texas, but has also been reported in Mississippi, Oklahoma and Louisiana. It is most common in the Pantanal region of Brazil and the Eastern Chaco region of Paraguay. It is the only member of the genus Jabiru. The name comes from the Tupi–Guaraní language and means "swollen neck".
The saddle-billed stork or saddlebill is a large wading bird in the stork family, Ciconiidae. It is a widespread species which is a resident breeder in sub-Saharan Africa from Sudan, Ethiopia and Kenya south to South Africa, and in The Gambia, Senegal, Côte d'Ivoire and Chad in west Africa. It is considered endangered in South Africa.
The greater adjutant is a member of the stork family, Ciconiidae. Its genus includes the lesser adjutant of Asia and the marabou stork of Africa. Once found widely across southern Asia and mainland southeast Asia, the greater adjutant is now restricted to a much smaller range with only three breeding populations; two in India, with the largest colony in Assam and a smaller one around Bhagalpur; and another breeding population in Cambodia. They disperse widely after the breeding season. This large stork has a massive wedge-shaped bill, a bare head and a distinctive neck pouch. During the day, it soars in thermals along with vultures with whom it shares the habit of scavenging. They feed mainly on carrion and offal; however, they are opportunistic and will sometimes prey on vertebrates. The English name is derived from their stiff "military" gait when walking on the ground. Large numbers once lived in Asia, but they have declined to the point of endangerment. The total population in 2008 was estimated at around a thousand individuals. In the 19th century, they were especially common in the city of Calcutta, where they were referred to as the "Calcutta adjutant" and included in the coat of arms for the city. Known locally as hargila and considered to be unclean birds, they were largely left undisturbed but sometimes hunted for the use of their meat in folk medicine. Valued as scavengers, they were once depicted in the logo of the Calcutta Municipal Corporation.
Ciconia is a genus of birds in the stork family. Six of the seven living species occur in the Old World, but the maguari stork has a South American range. In addition, fossils suggest that Ciconia storks were somewhat more common in the tropical Americas in prehistoric times.
The Oriental stork is a large, white bird with black-feathered wings in the stork family Ciconiidae.
Abdim's stork, also known as the white-bellied stork, is a stork belonging to the family Ciconiidae. It is the smallest species of stork, feeds mostly on insects, and is found widely in open habitats in Sub-Saharan Africa and in Yemen. The common name commemorates the Turkish Governor of Wadi Halfa in Sudan, Bey El-Arnaut Abdim (1780–1827).
Jabiru codorensis is an extinct species of stork related to the extant Jabiru. It lived in what is now Venezuela during the Pliocene period and appears to have been similar to its modern relative.