Mammuthus africanavus

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Mammuthus africanavus
Temporal range: Late Pliocene–Early Pleistocene
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Mammuthus africanavus plio tunisia.JPG
Molar found in Tunisia
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Proboscidea
Family: Elephantidae
Genus: Mammuthus
Species:
M. africanavus
Binomial name
Mammuthus africanavus
(Arambourg, 1952)
Synonyms

Elephas africanavus Arambourg, 1952

Mammuthus africanavus (literally, "African ancestor mammoth") is a species of mammoth known from remains spanning the Late Pliocene-Early Pleistocene found in Central and North Africa in the countries of Chad, Morocco, Tunisia and Algeria. It was originally described by Camille Arambourg in 1952 based on remains found around Lake Ichkeul in north Tunisia as a species of Elephas (which contains the living Asian elephant). [1] Some specimens from this sample may genuinely represent Elephas rather than Mammuthus, though the holotype has been argued to likely represent a true mammoth. [2] Some authors have argued that the species should be placed in Loxodonta (which contains living African elephants), reflecting the difficulty in distinguishing the teeth of early elephantids. [1] It is distinguished from the earlier Mammuthus subplanifrons by having a higher number of ridges/lamellae on the teeth (around 9-10 on the third molars), which display a greater parallelity, the molars being more hypsodont (high crowned), [1] with the molars having a greater amount of cementum and thinner enamel, and the molar plates exhibit closer spacing. [3]

The species is primarily known from dental remains. The skull and lower jaws of an old adult male mammoth, now apparently lost, referred to M. africanavus, has been reported from Garet et Tir, Algeria, dating to the Late Pliocene. The specimen, though somewhat damaged and distorted when described unambiguously represents a mammoth, with a tall skull over 1.45 metres (4.8 ft) in height with an especially short basicranium, and narrow palate. The specimen had twisted tusks at least 2.3 metres (7.5 ft) long, which were narrowly separated from each other in their elongate tusk sheaths/sockets/alveoli, but sharply diverged from each other following their emergence from the skull. The skull was described by Maglio (1973) as similar to that of the primitive Eurasian mammoth species Mammuthus meridionalis. [1]

Specimens intermediate between M. subplanifrons and M. africanavus have been reported from the Late Pliocene Hadar Formation, Ethiopia, dating to around 3.5 million years ago. [3] The oldest specimens of M. africanavus date to around 3.5-3 million years ago, while the youngest specimen dates to around 2.32-2 million years ago. [1]

At Aïn Boucherit in Algeria during the Early Pleistocene, the species lived alongside members of the genus Hippopotamus , rhinoceroses belonging to the genus Ceratotherium (which contains the living white rhinoceros), the suid (pig) Kolpochoerus, equines belonging to the genus Equus and Hipparion, the giant giraffid Sivatherium, the gomphothere Anancus, bovines, antelopes ( Gazella, Kobus, Damaliscus, Megalotragus, Parmularius, Oreonagor and Parantidorcas ) and canines ( Canis, Vulpes ). [4]

Related Research Articles

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A mammoth is any species of the extinct elephantid genus Mammuthus. They lived from the late Miocene epoch into the Holocene about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. Mammoths are distinguished from living elephants by their spirally twisted tusks and in at least some later species, the development of numerous adaptions to living in cold environments, including a thick layer of fur.

<span class="mw-page-title-main">Proboscidea</span> Order of elephant-like mammals

Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.

<span class="mw-page-title-main">Mastodon</span> Extinct genus of proboscideans

A mastodon is a member of the genus Mammut, which strictly defined, was endemic to North America and lived from the late Miocene to the early Holocene. Mastodons belong to the order Proboscidea, the same order as elephants and mammoths. Mammut is the type genus of the extinct family Mammutidae, which diverged from the ancestors of modern elephants at least 27-25 million years ago, during the Oligocene.

<span class="mw-page-title-main">Elephantidae</span> Family of mammals

Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.

<i>Elephas</i> Genus of mammals

Elephas is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, Elephas maximus. Several extinct species have been identified as belonging to the genus, extending back to the Pliocene or possibly late Miocene.

<i>Palaeoloxodon</i> Genus of extinct elephants

Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains some of the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.

<i>Stegodon</i> Genus of extinct proboscidean

Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon remained widespread in South, Southeast and East Asia until the end of the Pleistocene.

<span class="mw-page-title-main">Dwarf elephant</span> Prehistoric elephant species

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<i>Palaeoloxodon recki</i> Extinct species of elephant

Palaeoloxodon recki, often known by the synonym Elephas recki is an extinct species of elephant native to Africa and West Asia from the Pliocene or Early Pleistocene to the Middle Pleistocene. During most of its existence, the species represented the dominant elephant species in East Africa. The species is divided into five roughly chronologically successive subspecies. While the type and latest subspecies P. recki recki as well as the preceding P. recki ileretensis are widely accepted to be closely related to Eurasian Palaeoloxodon, the relationships of the other, chronologically earlier subspecies to P. recki recki and P. recki ileretensis are uncertain, with it being suggested they are unrelated and should be elevated to separate species.

<span class="mw-page-title-main">Columbian mammoth</span> Extinct species of mammoth that inhabited North America

The Columbian mammoth is an extinct species of mammoth that inhabited the Americas from southern Canada to Costa Rica during the Pleistocene epoch. The Columbian mammoth descended from Eurasian steppe mammoths that colonised North America during the Early Pleistocene around 1.5–1.3 million years ago, and later experienced hybridisation with the woolly mammoth lineage. The Columbian mammoth was among the last mammoth species, and the pygmy mammoths evolved from them on the Channel Islands of California. The closest extant relative of the Columbian and other mammoths is the Asian elephant.

<span class="mw-page-title-main">Straight-tusked elephant</span> Extinct species of elephant

The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. It was larger than any living elephant, with adult males suggested to reach 3.81–4.2 metres (12.5–13.8 ft) in shoulder height, and 11.3–15 tonnes in weight. Like modern elephants, the straight-tusked elephant lived in herds, flourishing during interglacial periods, when its range would extend as far north as Great Britain. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestral species of most dwarf elephants that inhabited islands in the Mediterranean.

<i>Anancus</i> Genus of proboscideans

Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.

<i>Mammuthus meridionalis</i> Extinct species of mammal

Mammuthus meridionalis, sometimes called the southern mammoth, is an extinct species of mammoth native to Eurasia, including Europe, during the Early Pleistocene, living from around 2.5 million years ago to 800,000 years ago.

<span class="mw-page-title-main">Steppe mammoth</span> Extinct species of mammal

Mammuthus trogontherii, sometimes called the steppe mammoth, is an extinct species of mammoth that ranged over most of northern Eurasia during the Early and Middle Pleistocene, approximately 1.7 million-200,000 years ago. One of the largest mammoth species, it evolved in East Asia during the Early Pleistocene, around 1.8 million years ago, before migrating into North America around 1.5 million years ago, and into Europe during the Early/Middle Pleistocene transition, around 1 to 0.7 million years ago. It was the ancestor of the woolly mammoth and Columbian mammoth of the later Pleistocene.

<i>Mammuthus lamarmorai</i> Extinct species of mammal

Mammuthus lamarmorai is a species of dwarf mammoth which lived during the late Middle and Late Pleistocene on the island of Sardinia in the Mediterranean. It has been estimated to have had a shoulder height of around 1.4 metres (4.6 ft). Remains have been found across the western part of the island.

Mammuthus subplanifrons is the oldest representative of the genus Mammuthus, known from around 6.2-3.75 million years ago during the late Miocene-early Pliocene in what is today South Africa and countries of East Africa, especially Ethiopia. They already presented some of the unique characteristics of mammoths like the spirally, twisting tusks.

<span class="mw-page-title-main">Woolly mammoth</span> Extinct species of mammoth from the Quaternary period

The woolly mammoth is an extinct species of mammoth that lived from the Middle Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with the African Mammuthus subplanifrons in the early Pliocene. The woolly mammoth began to diverge from the steppe mammoth about 800,000 years ago in Siberia. Its closest extant relative is the Asian elephant. The Columbian mammoth lived alongside the woolly mammoth in North America, and DNA studies show that the two hybridised with each other.

<i>Palaeoloxodon naumanni</i> Extinct species of elephant

Palaeoloxodon naumanni, occasionally called Naumann's elephant, is an extinct species belonging to the genus Palaeoloxodon found in the Japanese archipelago during the Middle to Late Pleistocene around 330,000 to 24,000 years ago. It is named after Heinrich Edmund Naumann who discovered the first fossils at Yokosuka, Kanagawa, Japan. Fossils attributed to P. naumanni are also known from China and Korea, though the status of these specimens is unresolved, and some authors regard them as belonging to separate species.

Mammuthus rumanus is a species of mammoth that lived during the Pliocene in Eurasia. It the oldest mammoth species known outside of Africa.

<i>Phanagoroloxodon</i> Extinct genus of mammals

Phanagoroloxodon is an extinct genus of elephant. It is known from one species Phanagoroloxodon mammontoides described from a partial skull found on the banks of the Psekups river in the northwestern Caucasus of Russia, of probable Late Pliocene-Early Pleistocene age. Phanagoroloxodon has been suggested to share a close common ancestry with Elephas, as well as mammoths, combining characteristics of both genera. Like the Asian elephant, the top of the skull has a groove running along the midline, while the tusks are suggested to be twisted, similar to those of mammoths. A 2020 PhD thesis by Steven Zhang suggested that Elephas recki brumpti from the Pliocene of East Africa should be subsumed into the species Elephas planifrons, known from the Late Pliocene-Early Pleistocene of the Indian subcontinent, and that this species should be placed as a second species of Phanagoroloxodon. However, these suggestions were rejected by Sanders (2023).

References

  1. 1 2 3 4 5 Sanders, William J. (2023-07-07). Evolution and Fossil Record of African Proboscidea (1 ed.). Boca Raton: CRC Press. pp. 245, 263–266. doi:10.1201/b20016. ISBN   978-1-315-11891-8.
  2. Markov, Georgi N. (October 2012). "Mammuthus rumanus, early mammoths, and migration out of Africa: Some interrelated problems". Quaternary International. 276–277: 23–26. doi:10.1016/j.quaint.2011.05.041.
  3. 1 2 Sanders, William J.; Haile-Selassie, Yohannes (June 2012). "A New Assemblage of Mid-Pliocene Proboscideans from the Woranso-Mille Area, Afar Region, Ethiopia: Taxonomic, Evolutionary, and Paleoecological Considerations". Journal of Mammalian Evolution. 19 (2): 105–128. doi:10.1007/s10914-011-9181-y. ISSN   1064-7554.
  4. Sahnouni, Mohamed; Van der Made, Jan; Everett, Melanie (June 2011). "Ecological background to Plio-Pleistocene hominin occupation in North Africa: the vertebrate faunas from Ain Boucherit, Ain Hanech and El-Kherba, and paleosol stable-carbon-isotope studies from El-Kherba, Algeria". Quaternary Science Reviews. 30 (11–12): 1303–1317. doi:10.1016/j.quascirev.2010.01.002.