Flies are insects of the order Diptera, the name being derived from the Greek δι- di- "two", and πτερόν pteron "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies, mosquitoes and others, although only about 125,000 species have been described.
The Tachinidae are a large and variable family of true flies within the insect order Diptera, with more than 8,200 known species and many more to be discovered. Over 1,300 species have been described in North America alone. Insects in this family commonly are called tachinid flies or simply tachinids. As far as is known, they all are protelean parasitoids, or occasionally parasites, of arthropods, usually other insects. The family is known from many habitats in all zoogeographical regions and is especially diverse in South America.
The Phoridae are a family of small, hump-backed flies resembling fruit flies. Phorid flies can often be identified by their escape habit of running rapidly across a surface rather than taking to the wing. This behaviour is a source of one of their alternate names, scuttle fly. Another vernacular name, coffin fly, refers to Conicera tibialis. About 4,000 species are known in 230 genera. The most well-known species is cosmopolitan Megaselia scalaris. At 0.4 mm in length, the world's smallest fly is the phorid Euryplatea nanaknihali.
The Sciaridae are a family of flies, commonly known as dark-winged fungus gnats. Commonly found in moist environments, they are known to be a pest of mushroom farms and are commonly found in household plant pots. This is one of the least studied of the large Diptera families, probably due to the small size of these insects and the similarity among species.
Delia flies are members of the Anthomyiidae family within the superfamily Muscoidae. The identification of different species of Delia can be very difficult for non-specialists as the diagnostic characteristics used for immature and/or female specimens may be inconsistent between species. Past taxonomic keys were not as comprehensive in their identification of Delia specimens; they were either too reliant on genetic characteristics, focused solely on a specific life stage, or were focused only on certain species. However current taxonomic keys aim to be more thorough by not only including morphological diagnostics for males, females, and immature specimens of various species, but also their genetic make-up or molecular barcode.
The fly Megaselia scalaris is a member of the order Diptera and the family Phoridae, and it is widely distributed in warm regions of the world. The family members are commonly known as the "humpbacked fly", the "coffin fly", and the "scuttle fly". The name "scuttle fly" derives from the jerky, short bursts of running, characteristic to the adult fly. The name "coffin fly" is due to their being found in coffins, digging six feet deep in order to reach buried corpses. It is one of the more common species found within the family Phoridae; more than 370 species have been identified within North America.
Rhagoletis cerasi is a species of fruit fly in the family Tephritidae.
The Metopininae are a subfamily of flies in the family Phoridae.
Anastrepha suspensa, known as the Caribbean fruit fly, the Greater Antillean fruit fly, guava fruit fly, or the Caribfly, is a species of tephritid fruit fly. As the names suggest, these flies feed on and develop in a variety of fruits, primarily in the Caribbean. They mainly infest mature to overripe fruits. While thought to have originated in Cuba, the Caribbean fruit fly can now also be found in Florida, Hispaniola, and Puerto Rico.
Megaselia nigra is a species of scuttle fly in the family Phoridae. Megaselia species are common pests of mushroom cultivation, attracted by the aroma of developing fungal mycelium. The larvae feed on the developing mycelium of the mushroom, ultimately damaging both the mycelium and gill tissues.
Howardula is a genus of nematode that infests the larvae of mushroom-feeding flies, beetles, and other insects. Various Howardula species and strains infest mushroom-feeding Drosophila, including Howardula aoronymphium and Howardula neocosmis. Howardula husseyi can infest the mushroom phorid Megaselia halterata.
Apocephalus paraponerae is a species of fly in the family Phoridae discovered by Borgmeier in 1958. This species is a parasitoid of the giant tropical ant Paraponera clavata and uses both visual and chemical cues to locate its host. A. paraponerae can locate fighting or injured ants through host-produced alarm pheromones. Female flies are attracted to the ant to feed and oviposit, while males are attracted to feed and locate females for mating. There is some evidence that suggests that A. paraponerae is a cryptic species complex of at least four genetically distinct species.
Pseudacteon tricuspis is a parasitic phorid fly that decapitates its host, the imported Solenopsis invicta fire ant. There are over 70 described species within the Pseudacteon genus, which parasitize a variety of ant species. However, P. tricuspis is very specific to its host ant and will not attack other native ant species, making it a good biological control against the fire ant. P. tricuspis was also introduced into the United States for this purpose. Aside from the United States, P. tricuspis has also been found in South America, Europe, and Asia. Female P. tricuspis deposit their eggs directly into the fire ant host. Deposition into the ant host determines the sex of the egg, which grows within the host until adulthood, killing and decapitating the host in the process. Interestingly, P. tricuspis has a male-biased sex ratio, where the males are smaller than the females.
The Metopininae is a tribe of flies in the family Phoridae.
Chonocephalus is a genus of flies in the family Phoridae.
Metopina is a genus of flies in the family Phoridae.
Microselia is a genus of flies in the family Phoridae.
Styletta is a genus of flies in the family Phoridae.
Aenigmatias is a genus of flies in the family Phoridae.
Myriophora is a genus of flies in the family Phoridae.
