Metriorhynchinae

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Metriorhynchines
Temporal range: Callovian–Valanginian
Metriorhynchus jaekeli.jpg
Thalattosuchus superciliosus skeleton, Senckenberg Museum of Frankfurt
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pseudosuchia
Clade: Crocodylomorpha
Suborder: Thalattosuchia
Family: Metriorhynchidae
Subfamily: Metriorhynchinae
Fitzinger, 1843
Genera

Metriorhynchinae is a subfamily of metriorhynchid crocodyliforms from the late Middle Jurassic to the Early Cretaceous (Callovian - Valanginian) of Europe, North America and South America. [1] [2] Named by Fitzinger, in 1843, it contains the metriorhynchids Maledictosuchus , Gracilineustes , Metriorhynchus , Cricosaurus and Rhacheosaurus . The last three taxa form a tribe within Metriorhynchinae, the Rhacheosaurini. Metriorhynchinae is one of two subfamilies of Metriorhynchidae, the other being Geosaurinae. [3]

Phylogeny

Metriorhynchinae is a stem-based taxon defined in 2009 as the most inclusive clade consisting of Metriorhynchus geoffroyii , but not Geosaurus giganteus . [4] Rhacheosaurini is a stem-based taxon and it was named and defined by Mark T. Young, Mark A. Bell and Stephen L. Brusatte in 2011 as the most inclusive clade including Rhacheosaurus gracilis , but not Metriorhynchus geoffroyii and Gracilineustes leedsi . [3] The cladogram below follows the topology from the analyses by Young, et al. 2020. [5] [6]

Metriorhynchinae 

Thalattosuchus superciliosus

Gracilineustes acutus

Gracilineustes leedsi

Maledictosuchus riclaensis

Maledictosuchus nuyivijanan

Metriorhynchus geoffroyii

 Rhacheosaurini 

Metriorhynchus palpebrosus

Cricosaurus saltillensis

Swiss rhacheosaurin

cf. Cricosaurus macrospondylus

Metriorhynchinae indet. (USNM 19640)

Rhacheosaurus gracilis

cf. Cricosaurus sp.

Cricosaurus araucanensis

Cricosaurus lithographicus

Cricosaurus schroederi

Cricosaurus vignaudi

Cricosaurus bambergensis

Cricosaurus suevicus

Cricosaurus elegans

Solnhofen Cricosaurus (C. rauhuti)

Related Research Articles

<span class="mw-page-title-main">Oxford Clay</span> Jurassic rock formation in southeast England

The Oxford Clay is a Jurassic marine sedimentary rock formation underlying much of southeast England, from as far west as Dorset and as far north as Yorkshire. The Oxford Clay Formation dates to the Jurassic, specifically, the Callovian and Oxfordian ages, and comprises two main facies. The lower facies comprises the Peterborough Member, a fossiliferous organic-rich mudstone. This facies and its rocks are commonly known as lower Oxford Clay. The upper facies comprises the middle Oxford Clay, the Stewartby Member, and the upper Oxford Clay, the Weymouth Member. The upper facies is a fossil poor assemblage of calcareous mudstones.

<i>Dakosaurus</i> Metriorhynchid reptile genus from the Late Jurassic / Early Cretaceous period

Dakosaurus is an extinct genus of crocodylomorph within the family Metriorhynchidae that lived during the Late Jurassic and Early Cretaceous. It was large, with teeth that were serrated and compressed lateromedially. The genus was established by Friedrich August von Quenstedt in 1856 for an isolated tooth named Geosaurus maximus by Theodor Plieninger in 1846. Dakosaurus was a carnivore that spent much, if not all, its life out at sea. The extent of its adaptation to a marine lifestyle means that it is most likely that it mated at sea, but since no eggs or nests have been discovered that have been referred to Dakosaurus, whether it gave birth to live young at sea like dolphins and ichthyosaurs or came ashore like turtles is not known. The name Dakosaurus means "biter lizard", and is derived from the Greek dakos ("biter") and σαῦρος -sauros ("lizard").

Aggiosaurus is an extinct genus of geosaurine metriorhynchid crocodyliform known from the Late Jurassic of Nice, southeastern France. It contains a single species, Aggiosaurus nicaeensis, which was named by H. Ambayrac in 1913.

<i>Metriorhynchus</i> Extinct genus of reptiles

Metriorhynchus is an extinct genus of marine crocodyliform that lived in the oceans during the Late Jurassic. The type species, M. brevirostris was named in 1829 as a species of Steneosaurus before being named as a separate genus by the German palaeontologist Christian von Meyer in 1832. The name Metriorhynchus means "Moderate snout", and is derived from the Greek Metrio- ("moderate") and -rhynchos ("snout").

<span class="mw-page-title-main">Metriorhynchidae</span> Extinct family of reptiles

Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period of Europe, North America and South America. The name Metriorhynchidae was coined by the Austrian zoologist Leopold Fitzinger in 1843. The group contains two subfamilies, the Metriorhynchinae and the Geosaurinae. They represent the most marine adapted of all archosaurs.

<i>Geosaurus</i> Extinct genus of reptiles

Geosaurus is an extinct genus of marine crocodyliform within the family Metriorhynchidae, that lived during the Late Jurassic and the Early Cretaceous. Geosaurus was a carnivore that spent much, if not all, its life out at sea. No Geosaurus eggs or nests have been discovered, so little is known of the reptile's lifecycle, unlike other large marine reptiles of the Mesozoic, such as plesiosaurs or ichthyosaurs which are known to give birth to live young out at sea. Where Geosaurus mated, whether on land or at sea, is currently unknown. The name Geosaurus means "Mother of Giants lizard", and is derived from the Greek Ge- and σαῦρος -sauros ("lizard"). The name Geosaurus was established by the French naturalist Georges Cuvier in 1824.

<i>Teleidosaurus</i> Genus of large reptiles

Teleidosaurus is an extinct genus of carnivorous metriorhynchoid crocodyliform from Middle Jurassic deposits of Normandy, France. The name Teleidosaurus means "Complete lizard", and is derived from the Greek Teleidos- ("complete") and σαῦρος -sauros ("lizard").

Purranisaurus is an extinct genus of marine crocodyliform from the Middle to Late Jurassic period of Chile and Vaca Muerta of Argentina. Rusconi originally regarded Purranisaurus potens to be a plesiosaur; however, Gasparini demonstrated that it was in fact a metriorhynchid crocodyliform, and that may be a junior synonym of Metriorhynchus. It was about 3–3.3 m (9.8–10.8 ft) long and weighed 80 kg (180 lb).

<i>Rhacheosaurus</i> Extinct genus of reptiles

Rhacheosaurus is an extinct genus of marine crocodyliform belonging to the family Metriorhynchidae. The genus was established by Christian Erich Hermann von Meyer in 1831 for skeletal remains from the Tithonian of Germany. It was a relatively small reptile, measuring between 1.38 and 1.57 m long and weighing 10 kg (22 lb).

<i>Suchodus</i> Extinct genus of reptiles

Suchodus is an extinct genus of marine crocodyliform from the Middle to Late Jurassic period of England and France. It measured between 3.3 and 4.7 m in total body length.

Cricosaurus is an extinct genus of marine crocodyliforms of the Late Jurassic. belonging to the family Metriorhynchidae. The genus was established by Johann Andreas Wagner in 1858 for three skulls from the Tithonian of Germany. The name Cricosaurus means "Ring lizard", and is derived from the Greek Krikos- ("ring") and σαῦρος -sauros ("lizard"). It was a relatively small reptile, with C. suevicus and C. araucanensis measuring 2 m (6.6 ft) and 3.2 m (10 ft) in total body length, respectively.

<i>Steneosaurus</i> Extinct genus of reptiles

Steneosaurus is a dubious genus of teleosaurid crocodyliform from the Middle or Late Jurassic of France. The genus has been used as a wastebasket taxon for thalattosuchian fossils for over two centuries, and almost all known historical species of teleosauroid have been included within it at one point. The genus has remained a wastebasket, with numerous species still included under the label ‘Steneosaurus’, many of which are unrelated to each other.

<i>Aeolodon</i> Extinct genus of reptiles

Aeolodon is an extinct genus of teleosaurid crocodyliform reptile from the Late Jurassic (Tithonian) of Germany and France that was initially named as a species of Crocodylus in 1814. Although previously synonymized with Steneosaurus, recent cladistic analysis considers it distantly related to the Steneosaurus type species and the type species is A. priscus, named in 1830 and described in 2020.

<i>Gracilineustes</i> Extinct genus of reptiles

Gracilineustes is an extinct genus of marine crocodyliform that lived in the oceans during the Middle to Late Jurassic. Gracilineustes was a carnivore that spent much, if not all, its life out at sea. It was a small reptile, with G. leedsi measuring 2.25–3.11 m (7.4–10.2 ft) long and G. acutus measuring 3.77 m (12.4 ft) long.

Eoneustes is an extinct genus of metriorhynchoid crocodyliform from Middle Jurassic deposits of France. Eoneustes was a carnivore that lived in the oceans and spent much, if not all, its life out at sea.

<span class="mw-page-title-main">Metriorhynchoidea</span> Extinct superfamily of reptiles

Metriorhynchoidea is an extinct superfamily of thalattosuchian crocodyliforms from the Early Jurassic to the Early Cretaceous of Europe, North America and South America. Metriorhynchids are fully aquatic crocodyliforms. Named by Fitzinger, in 1843, it contains the basal taxa like Teleidosaurus, Zoneait and Eoneustes and the family Metriorhynchidae. An unnamed taxon is known from Chile.

<i>Torvoneustes</i> Extinct genus of reptiles

Torvoneustes is an extinct genus of metriorhynchid thalattosuchian. It is known from skull and postcranial remains found in the Kimmeridge Clay Formation of Dorset and Wiltshire, England, the Virgula Marls of Switzerland and also from Oaxaca, Mexico . The holotype skull of the type species was initially assigned to the species Metriorhynchus superciliosus. Postcranial remains were later discovered from the same quarry as the skull, and then these specimens were recognised as belonging to a new species of Dakosaurus, as D. carpenteri. The species was named to honour Simon Carpenter, an amateur geologist from Frome in Somerset, who discovered the fossils.

<span class="mw-page-title-main">Geosaurinae</span> Extinct subfamily of reptiles

Geosaurinae is a subfamily of metriorhynchid crocodyliforms from the Middle Jurassic to the Early Cretaceous of Europe, North America and South America. Named by Richard Lydekker, in 1889, it contains the metriorhynchids Suchodus, Purranisaurus, Neptunidraco, Tyrannoneustes, Torvoneustes, Dakosaurus, Geosaurus and Plesiosuchus. The last four taxa form a tribe within Geosaurinae, the Geosaurini. Geosaurinae is one of two subfamilies of Metriorhynchidae, the other being Metriorhynchinae.

<i>Lemmysuchus</i> Extinct genus of reptiles

Lemmysuchus is a genus of machimosaurid thalattosuchian from the Middle Jurassic Callovian of England and France. Like many other teleosauroids from Europe, it has had a convoluted taxonomic history.

<i>Thalattosuchus</i> Extinct genus of reptiles

Thalattosuchus is an extinct genus of marine crocodyliform that lived in the oceans during the Middle to Late Jurassic. Thalattosuchus was named in 2020. Thalattosuchus was a carnivore that spent much, if not all, its life out at sea. No Thalattosuchus eggs or nests have been discovered, so little is known of the reptile's life cycle, unlike other large marine reptiles of the Mesozoic, such as plesiosaurs or ichthyosaurs which are known to have given birth to live young out at sea. Where Thalattosuchus mated, whether on land or at sea, is currently unknown. The name Thalattosuchus means "sea crocodile".

References

  1. Andrea Cau; Federico Fanti (2011). "The oldest known metriorhynchid crocodylian from the Middle Jurassic of North-eastern Italy: Neptunidraco ammoniticus gen. et sp. nov". Gondwana Research. 19 (2): 550–565. Bibcode:2011GondR..19..550C. doi:10.1016/j.gr.2010.07.007.
  2. Mark T. Young; Stephen L. Brusatte; Marcello Ruta; Marco Brandalise de Andrade (2010). "The evolution of Metriorhynchoidea (Mesoeucrocodylia, Thalattosuchia): an integrated approach using geometrics morphometrics, analysis of disparity and biomechanics". Zoological Journal of the Linnean Society. 158 (4): 801–859. doi: 10.1111/j.1096-3642.2009.00571.x .
  3. 1 2 Mark T. Young; Mark A. Bell; Stephen L. Brusatte (2011). "Craniofacial form and function in Metriorhynchidae (Crocodylomorpha: Thalattosuchia): modelling phenotypic evolution with maximum-likelihood methods". Biology Letters. 7 (6): 913–916. doi:10.1098/rsbl.2011.0357. PMC   3210659 . PMID   21543396.
  4. Mark T. Young; Marco Brandalise de Andrade (2009). "What is Geosaurus? Redescription of Geosaurus giganteus (Thalattosuchia: Metriorhynchidae) from the Upper Jurassic of Bayern, Germany". Zoological Journal of the Linnean Society. 157 (3): 551–585. doi: 10.1111/j.1096-3642.2009.00536.x .
  5. Young MT, Sachs S, Abel P, Foffa D, Herrera Y, Kitson JJ (2020). "Convergent evolution and possible constraint in the posterodorsal retraction of the external nares in pelagic crocodylomorphs". Zoological Journal of the Linnean Society. 189 (2): 494–520. doi: 10.1093/zoolinnean/zlaa021 . hdl: 11336/137182 .
  6. Young, M.T., Brignon, A., Sachs, S., Hornung J.J., Foffa, D., Kitson, J.J.N., Johnson, M.M., and Steel, L. 2020. Cutting the Gordian knot: a historical and taxonomic revision of the Jurassic crocodylomorph Metriorhynchus. Zoological Journal of the Linnean Society, zlaa092. https://doi.org/10.1093/zoolinnean/zlaa092.