Teleosauridae | |
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Platysuchus multiscrobiculatus , Holzmaden Germany | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauria |
Clade: | Pseudosuchia |
Clade: | Crocodylomorpha |
Clade: | Crocodyliformes |
Suborder: | † Thalattosuchia |
Parvorder: | † Neothalattosuchia |
Superfamily: | † Teleosauroidea |
Family: | † Teleosauridae Geoffroy, 1831 |
Subgroups | |
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Synonyms | |
Teleosauridae is a family of extinct typically marine crocodylomorphs similar to the modern gharial that lived during the Jurassic period. Teleosaurids were thalattosuchians closely related to the fully aquatic metriorhynchoids, but were less adapted to an open-ocean, pelagic lifestyle. The family was originally coined to include all the semi-aquatic (i.e. non-metriorhynchoid) thalattosuchians and was equivalent to the modern superfamily Teleosauroidea. However, as teleosauroid relationships and diversity was better studied in the 21st century, the division of teleosauroids into two distinct evolutionary lineages led to the establishment of Teleosauridae as a more restrictive family within the group, together with its sister family Machimosauridae.
Amongst teleosauroids, teleosaurids were generally smaller and less common than machimosaurids, suggesting the two families occupied different niches, similar to modern species of crocodilians. However, teleosaurids were more diverse than machimosaurids, with generalist coastal predators ( Mystriosaurus ), long-snouted marine piscivores ( Bathysuchus ), and potentially even long-snouted, semi-terrestrial predators ( Teleosaurus ). Additionally, teleosaurids occupied a wider range of habitats than machimosaurids, from semi-marine coasts and estuaries, the open-ocean, freshwater, and potentially even semi-terrestrial environments. [2]
Teleosauridae is phylogenetically defined by Mark T. Young as "the largest clade within Teleosauroidea containing Teleosaurus cadomensis but not Machimosaurus hugii [3] . Teleosauridae is split into two subfamilies, the Teleosaurinae and the Aeolodontinae, the former defined as the "the largest clade within Teleosauroidea containing Teleosaurus cadomensis, but not Aeolodon priscus and the latter defined as the "the largest clade within Teleosauroidea containing Aeolodon pricus, but not Teleosaurus cadomensis [3] .
Thalattosuchia |
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Teleosaurids were originally regarded as marine analogues to modern gharials, as they both typically share long, tubular snouts and narrow teeth. However, differences in the jaws, teeth, and skeleton of different teleosaurids suggest that they were more ecologically diverse than this. Earlier teleosaurids were coastal semi-aquatic generalists, while the two subfamilies were more specialised. Teleosaurines appear to have been semi-terrestrial, as they were more heavily armoured and had forward-facing nostrils. In contrast, aeolodontines have been found in deep marine waters and had reduced armour, implying that they were open water predators similar to metriorhynchoids (although the oldest aeolodontine, Mycterosuchus , appears to have been semi-terrestrial, similar to teleosaurines). [2] [4]
Definitive fossils of teleosaurids are restricted to Laurasia, with material found in Europe(England, France, Germany, Italy, Portugal, Russia and Switzerland) and Asia (China and Thailand, and possibly India). [2]
Machimosaurus is an extinct genus of machimosaurid crocodyliform from the Late Jurassic and Early Cretaceous. The type species, Machimosaurus hugii, was found in Switzerland. Other fossils have been found in England, France, Germany, Portugal, Switzerland and Tunisia. Machimosaurus rex is the largest named teleosauroid and thalattosuchian, with an estimated length of up to 7.15 m (23.5 ft). Machimosaurus is the largest known crocodyliform of the Jurassic.
Thalattosuchia is a clade of mostly marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Crocodilia and records from Thailand and China suggest that some members lived in freshwater. The clade contains two major subgroupings, the Teleosauroidea and Metriorhynchoidea. Teleosauroids are not greatly specialised for oceanic life, with back osteoderms similar to other crocodyliformes. Within Metriorhynchoidea, the Metriorhynchidae displayed extreme adaptions for life in the open ocean, including the transformation of limbs into flippers, the development of a tail fluke, and smooth, scaleless skin, and probably gave live birth, seemingly uniquely among archosaurs.
Teleosaurus is an extinct genus of teleosaurid crocodyliform found in the Middle Jurassic Calcaire de Caen Formation of France. It was approximately 3 metres (10 ft) in length. The holotype is MNHN AC 8746, a quarter of a skull and other associated postcranial remains, while other fragmentary specimens are known. The type species is T. cadomensis, but a second species, T. geoffroyi may also exist. It was previously considered a wastebasket taxon, with many other remains assigned to the genus.
Pelagosaurus is an extinct genus of thalattosuchian crocodyliform that lived during the Toarcian stage of the Lower Jurassic, around 183 Ma to 176 Ma, in shallow epicontinental seas that covered much of what is now Western Europe. The systematic taxonomy of Pelagosaurus has been fiercely disputed over the years, and was assigned to Thalattosuchia after its systematics within Teleosauridae were disputed. Pelagosaurus measured 2–3 m (6.6–9.8 ft) long.
Steneosaurus is a dubious genus of teleosaurid crocodyliform from the Middle or Late Jurassic of France. The genus has been used as a wastebasket taxon for thalattosuchian fossils for over two centuries, and almost all known historical species of teleosauroid have been included within it at one point. The genus has remained a wastebasket, with numerous species still included under the label ‘Steneosaurus’, many of which are unrelated to each other.
Aeolodon is an extinct genus of teleosaurid crocodyliform reptile from the Late Jurassic (Tithonian) of Germany and France that was initially named as a species of Crocodylus in 1814. Although previously synonymized with Steneosaurus, recent cladistic analysis considers it distantly related to the Steneosaurus type species and the type species is A. priscus, named in 1830 and described in 2020.
Mycterosuchus is an extinct genus of teleosaurid crocodyliform from the Middle Jurassic (Callovian) of England. Although previously synonymized with Steneosaurus, recent cladistic analysis considers it distantly related to the Steneosaurus type species.
Sericodon is an extinct genus of teleosaurid crocodyliform from the Late Jurassic (Tithonian) of Germany and Switzerland. The genus contains a single species, S. jugleri. Sericodon was placed in 'Clade T' (Aeolodontinae) and was found to be the sister taxon to Bathysuchus, another teleosaurid.
Metriorhynchoidea is an extinct superfamily of thalattosuchian crocodyliforms from the Early Jurassic to the Early Cretaceous of Europe, North America and South America. Metriorhynchids are fully aquatic crocodyliforms. Named by Fitzinger, in 1843, it contains the basal taxa like Teleidosaurus, Zoneait and Eoneustes and the family Metriorhynchidae. An unnamed taxon is known from Chile.
Torvoneustes is an extinct genus of metriorhynchid thalattosuchian. It is known from skull and postcranial remains found in the Kimmeridge Clay Formation of Dorset and Wiltshire, England, the Virgula Marls of Switzerland and also from Oaxaca, Mexico . The holotype skull of the type species was initially assigned to the species Metriorhynchus superciliosus. Postcranial remains were later discovered from the same quarry as the skull, and then these specimens were recognised as belonging to a new species of Dakosaurus, as D. carpenteri. The species was named to honour Simon Carpenter, an amateur geologist from Frome in Somerset, who discovered the fossils.
Macrospondylus is an extinct genus of machimosaurid teleosauroid crocodyliform from the Early Jurassic (Toarcian) of Europe. Fossils are known from the Posidonia Shale of Germany, the Whitby Mudstone of the United Kingdom, and the "schistes bitumineux" of Luxembourg.
Bathysuchus is an extinct genus of teleosaurid thalattosuchian from Late Jurassic (Kimmeridgian) deep water marine deposits in England and France. Bathysuchus displays features that suggest it was more pelagic than other teleosaurids, including smoother skull bones and reduced armour plating, similar to the fully marine metriorhynchids. This was possibly an adaptation to rising sea levels during the Kimmeridgian, as its earlier relatives such as Teleosaurus were suited for shallow coasts and lagoon environments.
Lemmysuchus is a genus of machimosaurid thalattosuchian from the Middle Jurassic Callovian of England and France. Like many other teleosauroids from Europe, it has had a convoluted taxonomic history.
Indosinosuchus is a genus of teleosaurid neosuchian that lived during the Late Jurassic or Early Cretaceous in what is now Thailand. It contains two species, the type species I. potamosiamensis and I. kalasinensis, both recovered from the lower Phu Kradung Formation. It is unique among teleosauroids as it is the only named genus known from a freshwater environment, while most other members of the group are marine. Indosinosuchus is placed in the family Teleosauridae, but has a relatively robust skull that bears resemblance to members of the Machimosauridae. Biomechanical analysis of its mandible and teeth suggest that it would have had a substantial bite force comparable to animals like Lemmysuchus. The two Indosinosuchus species however differ in the speed at which they could open and close their jaws, impacting their respective ecology and possibly explaining how they coexisted in the same environment. All known specimens of this genus were recovered from a single locality, which has been interpreted as a mass death site, possibly caused by a drought or flash flood. The precise age of Indosinosuchus is unclear, as the vertebrate fossils of the Phu Kradung Formation support a Late Jurassic age, while palynological data suggests an Early Cretaceous (Berriasian) age.
Teleosauroidea is an extinct superfamily of thalattosuchian crocodyliforms living from the Early Jurassic to the Early Cretaceous. It is phylogenetically defined by Mark T. Young and colleagues in 2024 as "the largest clade within Thalatttosuchia containing Teleosaurus cadomensis, but not Metriorhynchus brevirostris. This group contains two main families, the more piscivorous and gracile Teleosauridae and the more macropredatory and robust Machimosauridae.
Plagiophthalmosuchus is a genus of teleosauroid, known form the Early Jurassic Whitby Mudstone Formation of Whitby, Yorkshire, UK, and Dudelange, Luxembourg. The type species, P. gracilirostris, was originally named as a species of Teleosaurus in 1836, but then it was moved to Steneosaurus in 1961, but it was again moved to its own genus in 2020.
Neosteneosaurus is a genus of machimosaurid, known from the Middle Jurassic Oxford Clay of the UK, and Marnes de Dives, France. The type species, N. edwardsi, was originally named as a species of Steneosaurus in 1868, but was moved to its own genus in 2020. Steneosaurus durobrivensis and Steneosaurus hulkei are considered junior synonyms.
Machimosauridae is an extinct family of teleosauroid thalattosuchian crocodyliforms. The family was first identified in 2016, when fossils of teleosauroid thalattosuchians, including an indeterminate close relative of Lemmysuchus and Machimosaurus, were described from the Middle Jurassic (Bathonian) of Morocco. The family was largely expanded in 2020 when the systematics of Teleosauroidea were re-reviewed. Members of this family generally were larger than the teleosaurids.
Clovesuurdameredeor is an extinct genus of machimosaurid teleosauroid from the Bathonian Cornbrash Formation of England.
Seldsienean in an extinct genus of machimosaurid thalattosuchian from the Middle Jurassic of England and France. It is known from the Calcaire de Caen and the Cornbrash Formation.