Enaliosuchus

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Enaliosuchus
Temporal range: Early Cretaceous,
~140–132  Ma
Enaliosuchus BW.jpg
Restoration of E. macrospondylus
Scientific classification
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Enaliosuchus

Koken, 1883 [1]
Type species
Enaliosuchus macrospondylus
Koken, 1883 [1]
Synonyms

Enaliosuchus is a dubious [2] genus of extinct marine crocodyliform within the family Metriorhynchidae that lived during the Valanginian stage of the Early Cretaceous. It is known from fossil remains found in France and Germany and it was first described in 1883. [1]

Contents

Only the type species is known, which is E. macrospondylus. [1]

Etymology

The name Enaliosuchus means "Marine crocodile", and is derived from the Greek Enalios- ("marine") and -suchos ("crocodile"). [1]

Discovery and naming

Vertebra from the holotype of E. macrospondylus Enaliosuchus vertebra holotype.png
Vertebra from the holotype of E. macrospondylus

The holotype, specimen MB.R.1943.1–16, consists of an atlas-axis complex, three post-axial cervical vertebrae, several dorsal vertebrae, a caudal vertebra, an incomplete femur and a fragmentary sacral rib, and it was discovered in the Stadthagen Formation in Germany. The species Enaliosuchus macrospondylus was then named for this specimen by Koken (1883). [1]

Isolated teeth were subsequently found across Germany and have been assigned to Enaliosuchus, [2] and a partial specimen (specimen RNGD 990201) discovered in the Campylotoxus Zone of France was assigned to Enaliosuchus macrospondylus by Hua et al. (2000). [3]

Description

Enaliosuchus was a carnivore that spent much, if not all, its life out at sea. [3] No Enaliosuchus eggs or nest have been discovered, so little is known of the reptile's lifecycle, unlike other large marine reptiles of the Mesozoic, such as plesiosaurs or ichthyosaurs which are known to give birth to live young out at sea. Where Enaliosuchus mated, whether on land or at sea, is currently unknown. [3]

Species

The species within Enaliosuchus include :

Recent phylogenetic analyses found the genus Enaliosuchus to be monophyletic, but also to be nested within the genus Geosaurus or Cricosaurus . [4] [5] Sachs et al. (2020) hypothesised that the two species within the genus Enaliosuchus were not monophyletic. They hypothesised that E. schroederi belonged to a new, undescribed, genus. [2]

In 2024, Enaliosuchus schroederi, from Germany, [6] which was previously believed to likely be a junior synonym of E. macrospondylus, [3] was moved to the new genus Enalioetes in 2024. [7]

Related Research Articles

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Dakosaurus is an extinct genus of crocodylomorph within the family Metriorhynchidae that lived during the Late Jurassic and Early Cretaceous. It was large, with teeth that were serrated and compressed lateromedially. The genus was established by Friedrich August von Quenstedt in 1856 for an isolated tooth named Geosaurus maximus by Theodor Plieninger in 1846. Dakosaurus was a carnivore that spent much, if not all, its life out at sea. The extent of its adaptation to a marine lifestyle means that it is most likely that it mated at sea, but since no eggs or nests have been discovered that have been referred to Dakosaurus, whether it gave birth to live young at sea like dolphins and ichthyosaurs or came ashore like turtles is not known yet. The name Dakosaurus means "biter lizard", and is derived from the Greek dakos ("biter") and σαῦρος -sauros ("lizard").

<i>Machimosaurus</i> Genus of reptiles

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<i>Metriorhynchus</i> Extinct genus of reptiles

Metriorhynchus is an extinct genus of marine crocodyliform that lived in the oceans during the Late Jurassic. The type species, M. brevirostris was named in 1829 as a species of Steneosaurus before being named as a separate genus by the German palaeontologist Christian von Meyer in 1832. The name Metriorhynchus means "Moderate snout", and is derived from the Greek Metrio- ("moderate") and -rhynchos ("snout").

<span class="mw-page-title-main">Metriorhynchidae</span> Extinct family of reptiles

Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period of Europe, North America and South America. The name Metriorhynchidae was coined by the Austrian zoologist Leopold Fitzinger in 1843. The group contains two subfamilies, the Metriorhynchinae and the Geosaurinae. They represent the most marine adapted of all archosaurs.

<i>Geosaurus</i> Extinct genus of reptiles

Geosaurus is an extinct genus of marine crocodyliform within the family Metriorhynchidae, that lived during the Late Jurassic and the Early Cretaceous. Geosaurus was a carnivore that spent much, if not all, its life out at sea. No Geosaurus eggs or nests have been discovered, so little is known of the reptile's lifecycle, unlike other large marine reptiles of the Mesozoic, such as plesiosaurs or ichthyosaurs which are known to give birth to live young out at sea. Where Geosaurus mated, whether on land or at sea, is currently unknown. The name Geosaurus means "Mother of Giants lizard", and is derived from the Greek Ge- and σαῦρος -sauros ("lizard"). The name Geosaurus was established by the French naturalist Georges Cuvier in 1824.

<i>Teleidosaurus</i> Genus of large reptiles

Teleidosaurus is an extinct genus of carnivorous metriorhynchoid crocodyliform from Middle Jurassic deposits of Normandy, France. The name Teleidosaurus means "Complete lizard", and is derived from the Greek Teleidos- ("complete") and σαῦρος -sauros ("lizard").

<span class="mw-page-title-main">Thalattosuchia</span> Clade of marine crocodylomorphs

Thalattosuchia is a clade of mostly marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Crocodilia and records from Thailand and China suggest that some members lived in freshwater. The clade contains two major subgroupings, the Teleosauroidea and Metriorhynchoidea. Teleosauroids are not greatly specialised for oceanic life, with back osteoderms similar to other crocodyliformes. Within Metriorhynchoidea, the Metriorhynchidae displayed extreme adaptions for life in the open ocean, including the transformation of limbs into flippers, the development of a tail fluke, and smooth, scaleless skin, and probably gave live birth, seemingly uniquely among archosaurs.

Purranisaurus is an extinct genus of marine crocodyliform from the Middle to Late Jurassic period of Chile and Vaca Muerta of Argentina. Rusconi originally regarded Purranisaurus potens to be a plesiosaur; however, Gasparini demonstrated that it was in fact a metriorhynchid crocodyliform, and that may be a junior synonym of Metriorhynchus. It was about 3.3 m (11 ft) long.

<i>Rhacheosaurus</i> Extinct genus of reptiles

Rhacheosaurus is an extinct genus of marine crocodyliform belonging to the family Metriorhynchidae. The genus was established by Christian Erich Hermann von Meyer in 1831 for skeletal remains from the Tithonian of Germany. It was a relatively small reptile, measuring between 1.39 and 1.57 m long.

<i>Cricosaurus</i> Extinct genus of reptiles

Cricosaurus is an extinct genus of marine crocodyliforms of the Late Jurassic. belonging to the family Metriorhynchidae. The genus was established by Johann Andreas Wagner in 1858 for three skulls from the Tithonian of Germany. The name Cricosaurus means "Ring lizard", and is derived from the Greek Krikos- ("ring") and σαῦρος -sauros ("lizard"). It was a relatively small reptile, with C. suevicus and C. araucanensis measuring 2 m (6.6 ft) and 3.2 m (10 ft) in total body length, respectively.

<i>Brancasaurus</i> Extinct genus of reptiles

Brancasaurus is a genus of plesiosaur which lived in a freshwater lake in the Early Cretaceous of what is now North Rhine-Westphalia, Germany. With a long neck possessing vertebrae bearing distinctively-shaped "shark fin"-shaped neural spines, and a relatively small and pointed head, Brancasaurus is superficially similar to Elasmosaurus, albeit smaller in size at 3.26 metres (10.7 ft) in length as a subadult.

<span class="mw-page-title-main">Metriorhynchoidea</span> Extinct superfamily of reptiles

Metriorhynchoidea is an extinct superfamily of thalattosuchian crocodyliforms from the Early Jurassic to the Early Cretaceous of Europe, North America and South America. Metriorhynchids are fully aquatic crocodyliforms. Named by Fitzinger, in 1843, it contains the basal taxa like Teleidosaurus, Zoneait and Eoneustes and the family Metriorhynchidae. An unnamed taxon is known from Chile.

<i>Torvoneustes</i> Extinct genus of reptiles

Torvoneustes is an extinct genus of metriorhynchid thalattosuchian. It is known from skull and postcranial remains found in the Kimmeridge Clay Formation of Dorset and Wiltshire, England, the Virgula Marls of Switzerland and also from Oaxaca, Mexico . The holotype skull of the type species was initially assigned to the species Metriorhynchus superciliosus. Postcranial remains were later discovered from the same quarry as the skull, and then these specimens were recognised as belonging to a new species of Dakosaurus, as D. carpenteri. The species was named to honour Simon Carpenter, an amateur geologist from Frome in Somerset, who discovered the fossils.

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<span class="mw-page-title-main">Geosaurinae</span> Extinct subfamily of reptiles

Geosaurinae is a subfamily of metriorhynchid crocodyliforms from the Middle Jurassic to the Early Cretaceous of Europe, North America and South America. Named by Richard Lydekker, in 1889, it contains the metriorhynchids Suchodus, Purranisaurus, Neptunidraco, Tyrannoneustes, Torvoneustes, Dakosaurus, Geosaurus and Plesiosuchus. The last four taxa form a tribe within Geosaurinae, the Geosaurini. Geosaurinae is one of two subfamilies of Metriorhynchidae, the other being Metriorhynchinae.

<span class="mw-page-title-main">Metriorhynchinae</span> Extinct subfamily of reptiles

Metriorhynchinae is a subfamily of metriorhynchid crocodyliforms from the late Middle Jurassic to the Early Cretaceous of Europe, North America and South America. Named by Fitzinger, in 1843, it contains the metriorhynchids Maledictosuchus, Gracilineustes, Metriorhynchus, Cricosaurus and Rhacheosaurus. The last three taxa form a tribe within Metriorhynchinae, the Rhacheosaurini. Metriorhynchinae is one of two subfamilies of Metriorhynchidae, the other being Geosaurinae.

<i>Maledictosuchus</i> Extinct genus of reptiles

Maledictosuchus is an extinct genus of marine crocodyliform belonging to the family Metriorhynchidae. It is the most basal member of the Rhacheosaurini Tribe with a relatively short body length, measuring 2.95 m (9.7 ft) based on the type specimen.

<i>Zoneait</i> Extinct genus of reptiles

Zoneait is an extinct genus of thalattosuchian crocodylomorph known from a single species, Zoneait nargorum, from the Middle Jurassic Snowshoe Formation of Oregon.

Enalioetes is an extinct genus of metriorhynchid thalattosuchian from the Early Cretaceous Stadthagen Formation of Germany. The type species is E. schroederi.

The Stadthagen Formation is a geological formation located in Hanover, Germany, dating to the Early Cretaceous (Valanginian). The formation contains fossils of reptiles and fish, suggesting that the formation was once a shallow marine setting.

References

  1. 1 2 3 4 5 6 7 Koken E. (1883). Die reptilian der norddeutschen unteren Kreide. Zeitschrift deutschen Geologischen Gesellschaft35: 735-827.
  2. 1 2 3 Sachs, Sven; Young, Mark T.; Hornung, Jahn J. (2020). "The enigma of Enaliosuchus, and a reassessment of the Lower Cretaceous fossil record of Metriorhynchidae". Cretaceous Research. 114: 104479. doi:10.1016/j.cretres.2020.104479. hdl: 20.500.11820/c52d1d56-1bf3-4aae-b2e1-38c85eed44fa . ISSN   0195-6671. S2CID   218996914.
  3. 1 2 3 4 5 Hua, S., Vignaud, P., Atrops, F. & Clément, A. (2000). Enaliosuchus macrospondylus Koken, 1883 (Crocodylia, Metriorhynchidae) du Valanginien de Barret-le-Bas (Hautes Alpes, France): un cas unique de remontée des narines externes parmi les crocodiliens. Geobios33: 467-474.
  4. Young M. T. (2007). The evolution and interrelationships of Metriorhynchidae (Crocodyliformes, Thalattosuchia). Journal of Vertebrate Paleontology27 (3): 170A.
  5. Wilkinson L. E, Young M. T, Benton M. J. (2008). A new metriorhynchid crocodilian (Mesoeucrocodylia: Thalattosuchia) from the Kimmeridgian (Upper Jurassic) of Wiltshire, UK. Palaeontology51 (6): 1307-1333.
  6. "Kuhn, O. (1936). Crocodilia. Fossilium Catalogus I: Animalia 75. Gracenhage: Junk, 114pp.
  7. Sachs, S.; Young, M.T.; Hornung, J.J.; Cowgill, T.; Schwab, J.A.; Brusatte, S.L. (2024). "A new genus of metriorhynchid crocodylomorph from the Lower Cretaceous of Germany". Journal of Systematic Palaeontology. 22 (1). doi:10.1080/14772019.2024.2359946.