Ogygiocarella Temporal range: | |
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Ogygiocarella debuchii Brongniart, 1822, from the Meadowtown Formation, Llandeilo Group, Middleton Series, Abereiddian Stage, Meadowtown, Shropshire. Length 8.5 cm. | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | † Trilobita |
Order: | † Asaphida |
Family: | † Asaphidae |
Genus: | † Ogygiocarella Harrington & Leanza, 1957 |
Type species | |
Asaphus debuchii | |
Species | |
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Ogygiocarella Brongniart, 1822, [1] is a genus of asaphid trilobites. It occurred during the Middle Ordovician. [2] [3]
The generic name, Ogygiocarella, refers to "Ogygia", the seventh daughter of Amphion and Niobe, which name is combined with "-care-", from the Greek "akares", meaning short, and finally "-ella", the diminutive form.[ citation needed ]
Ogygiocarella is arguably the first trilobite that was scientifically described. Rev. Edward Lhwyd [4] published in 1698 in The Philosophical Transactions of the Royal Society, the oldest scientific journal in the English language, part of his letter “Concerning Several Regularly Figured Stones Lately Found by Him", that was accompanied by a page of etchings of fossils. [5] One of his etchings figured a trilobite he found near Llandeilo, probably on the grounds of Lord Dynefor's castle, he described as “… the skeleton of some flat Fish …". [6]
Ogygiocarella has a very flat calcified dorsal exoskeleton, with an oval outline (about 1½× longer than wide). The headshield (or cephalon) is 2½× wider than long, and of equal size as the tailshield (or pygidium), a state called isopygeous. Both are semicircular. The backcorners of the cephalon end in so-called genal spines that stick backwards approximately to the 6th thorax segment. The central raised area of the cephalon (or glabella) has 4 pairs of furrows and expands in front of the eyes. The eyes are small, close to the glabella and in the rear half of the cephalon. The dorsal facial sutures, that split when moulding, arch from the front of the eye and pass in front of the glabella. The suture follows the top of the visual surface, as in all trilobites, and from there cut backwards and outwards at approximately 45° to reach the posterior margin of the cephalon approximately halfway between the glabella and the lateral border. As usual for members of the family Asaphidae, it has 8 articulating segments in the middle part of the body (or thorax). The axis is about half as wide as the ribs (or pleurae) to each of the sides. In the thorax the axis tapers slightly towards the back. In the pygidium the axis tapers stronger and terminates a short distance from the margin in a roundish endpiece. It has 10-14 ribs on each side of the axis, which fade-out in the border that is as wide as the axis. [7]
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.
Dalmanites is a genus of trilobite in the order Phacopida. They lived from the Late Ordovician to Middle Devonian.
Asaphus is a genus of trilobites that is known from the Lower and Middle Ordovician of northwestern Europe.
Dikelocephalus is a genus of very large trilobites of up to 50 cm (20 in) long, that lived during the last 3 million years of the Cambrian (Sunwaptan). Their fossils are commonly found as disarticulated sclerites, in the upper Mississippi Valley and in Canada (Alberta). The exoskeleton is rounded anteriorly, with the thorax and sides of the tailshield slightly tapering to about ⅔× of the width across the base of the spines at the back of the headshield. At the side corners of the pygidium there may be triangular or hooked spines, pointing backwards, while between the spines the posterior margin is at a 30-75° angle with the lateral margin, gently convex or nearly straight. If pygidial spines are lacking, the margin is gradually rounded. The thorax has 12 segments.
Meniscuchus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the Botomian stage, which lasted from approximately 522 to 516 million years ago. This faunal stage was part of the Cambrian Period. Meniscuchus has been found in the USA, Canada, Russia and Australia.
Meteoraspis is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae. The various species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Meteoraspis are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as M. nevensis from Victoria Land, Antarctica.
Ptychoparia is a genus of ptychopariid trilobites, and is the type genus of the family Ptychopariidae, and the order Ptychopariida.
Conocoryphe is a genus of primarily eyeless trilobites belonging to the family Conocoryphidae. They lived during the Middle Cambrian period, about 505 million years ago. These arthropods lived on the sea bottom (epifaunal) and lived off dead particulate organic matter.
Odontochile is a genus of trilobites in the order Phacopida, family Dalmanitidae.
Litometopus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the Botomian stage.
Galbagnostus is an extinct genus of agnostid trilobite. It lived during the Lower and Middle Ordovician.
Sphaeragnostus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It can be recognized by having two thorax segments, a totally effaced headshield, while the tailshield although effaced, has a clear furrow parallel to its border, and a short, convex, subcircular axis. It lived during the Ordovician.
Thoracocare is a minute to very small trilobite, that lived during part of the Middle Cambrian in what are today the states of Idaho, Nevada and Utah. It is the only trilobite known with just two thorax segments outside most members of the Agnostida order. It can be distinguished from Agnostida by the very wide subquadrate glabella, parallel-side or widening forward in the largest specimen, with the full front side touching the border. Two species are known, one, T. idahoensis, only from pygidia.
Geragnostus is a genus of very small agnostid trilobites whose fossils are found Ordovician-aged marine strata from Eurasia, North America and Argentina.
Cnemidopyge is a genus of trilobites that lived during the Ordovician. Like all Raphiophorids it is blind, with a cephalon that is subtriangular to subsemicircular, carrying genal spines and a forward directed rapier-like spine on the central raised area, with the front of the glabella inflated and the natural fracture lines of the cephalon coinciding with its margin. It may be easily distinguished from other raphiophorids by the rectangular thorax with 6 segments, where other genera have a different number of segments and segments change in width over the length of the thorax. Uniquely in this genus, the inner pleural region of the frontal segment is enlarged. Also the axis and pleural fields of the pygidium are strongly segmented.
Cedaria is a small, rather flat trilobite with an oval outline, a headshield and tailshield of approximately the same size, 7 articulating segments in the middle part of the body and spines at the back edges of the headshield that reach halflength of the body. Cedaria lived during the early part of the Upper Cambrian (Dresbachian), and is especially abundant in the Weeks Formation.
Orygmaspis is a genus of asaphid trilobite with an inverted egg-shaped outline, a wide headshield, small eyes, long genal spines, 12 spined thorax segments and a small, short tailshield, with four pairs of spines. It lived during the Upper Cambrian in what are today Canada and the United States.
Placoparia is a genus of trilobites of average size that lived during the late Lower to the early Upper Ordovician on the paleocontinents Gondwana, Avalonia and Laurentia, now the Czech Republic, France, Germany, Morocco, Portugal, Spain and Wales. Its headshield is semi-circular to rectangular with rounded frontal corners. It lacks eyes, but eye ridges are present. The fact that the facial sutures are opisthoparian is an exception in the otherwise proparian Cheirurina. The thorax has 11 or 12 segments, with the axis slightly wider than the ribs to its sides. The tips of the pleurae are free, which resembles an old-fashion central heating radiator. The axis in the small tailshield consists of four rings and a minute endpiece. The four pleurae end in spatulate spines that fit to corresponding indentations in the cephalon.
Tricrepicephalus is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae with species of average size. Its species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Tricrepicephalus are widespread in Late Cambrian deposits in North America, but is also known from one location in South-America. Tricrepicephalus has an inverted egg-shaped exoskeleton, with three characteristic pits in the fold that parallels the margin of the headshield just in front of the central raised area. The articulating middle part of the body has 12 segments and the tailshield carries two long, tubular, curved pygidial spines that are reminiscent of earwig's pincers that rise backwards from the plain of the body at approximately 30°.