Babblers | |
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Pin-striped tit-babbler, (Mixornis gularis) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Passeriformes |
Superfamily: | Sylvioidea |
Family: | Timaliidae Vigors & Horsfield, 1827 |
Genera | |
See article text |
The Old World babblers or Timaliidae, are a family of mostly Old World passerine birds. They are rather diverse in size and coloration, but are characterised by soft, fluffy plumage. These are birds of tropical areas, with the greatest variety in Southeast Asia and the Indian subcontinent. The timaliids are one of two unrelated groups of birds known as babblers, the other being the Australasian babblers of the family Pomatostomidae (also known as pseudo-babblers).
Morphological diversity is rather high; most species resemble "warblers", jays or thrushes. This group is among those Old World bird families with the highest number of species still being discovered.
Timaliids are small to medium birds. They have strong legs, and many are quite terrestrial. They typically have generalised bills, similar to those of a thrush or warbler, except for the scimitar babblers which, as their name implies, have strongly decurved bills. Most have predominantly brown plumage, with minimal difference between the sexes, but many more brightly coloured species also exist. [1]
This group is not strongly migratory, and most species have short rounded wings, and a weak flight. They live in lightly wooded or scrubland environments, ranging from swamp to near-desert. They are primarily insectivorous, although many will also take berries, and the larger species will even eat small lizards and other vertebrates. [1]
Typical babblers live in communities of around a dozen birds, jointly defending a territory. Many even breed communally, with a dominant pair building a nest, and the remainder helping to defend and rear their young. Young males remain with the group, while females move away to find a new group, and thus avoid inbreeding. They make nests from twigs, and hide them in dense vegetation. [1]
The systematics of Old World babblers have long been contested. During much of the 20th century, the family was used as a "wastebin taxon" for numerous hard-to-place Old World songbirds (such as Picathartidae and Pnoepygidae, as well as the New World species the wrentit). The German ornithologist Ernst Hartert summarized this attitude with the statement that, in the passerines: "Was man nicht unterbringen kann, sieht man als Timalien an." (What one can't place systematically is considered an Old World babbler). [2]
The most obviously misplaced taxa were removed piecemeal towards the end of the last century. Since then, with the aid of DNA sequence data, it has been confirmed that even the remaining group is not monophyletic. Analysis of mtDNA cytochrome b and 12S/16S rRNA data (Cibois 2003a) spread the Timaliidae that were studied across what essentially was a badly resolved polytomy with Old World warblers and white-eyes. As the typical warblers (genus Sylvia ) grouped with some presumed timaliids (such as the fulvettas), it was suggested that some Sylviidae should be moved to the Timaliidae.
The phylogenetic relationships between Timaliidae and other families was determined in a molecular phylogenetic study by Tianlong Cai and collaborators that was published in 2019. It is shown in the cladogram below: [3] [4]
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The cladogram below shows the relationships between the genera. These were determined in the same study by Cai and collaborators. [3] [4]
Timaliidae |
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The family as currently constituted includes 58 species divided into the following ten genera: [4]
Image | Genus | Living species |
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Erythrogenys Baker, 1930 – scimitar-babblers |
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Pomatorhinus Horsfield, 1821 – scimitar-babblers |
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Spelaeornis David & Oustalet, 1877 – typical wren-babblers |
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Stachyris Hodgson, 1844 |
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Cyanoderma Salvadori, 1874 [5] |
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Dumetia Blyth, 1852 |
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Mixornis Blyth, 1842 – tit-babblers |
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Macronus Jardine & Selby, 1835 – tit-babblers |
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Timalia Horsfield, 1821 |
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Melanocichla Sharpe, 1883 |
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Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. They are not closely related to the New World warblers. The family held over 400 species in over 70 genera, and were the source of much taxonomic confusion. Two families were split out initially, the cisticolas into Cisticolidae and the kinglets into Regulidae. In the past ten years they have been the subject of much research and many species are now placed into other families, including the Acrocephalidae, Cettiidae, Phylloscopidae, and Megaluridae. In addition some species have been moved into existing families or have not yet had their placement fully resolved. A smaller number of warblers, together with some babblers formerly placed in the family Timaliidae and the parrotbills, are retained in a much smaller family Sylviidae.
Sylviidae is a family of passerine birds that includes the typical warblers and a number of babblers formerly placed within the Old World babbler family. They are found in Eurasia and Africa.
The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World, with the exception of several vagrants and two species, bluethroat and northern wheatear, found also in North America. These are mainly small arboreal insectivores, many of which, as the name implies, take their prey on the wing. The family is relatively large and includes 351 species which are divided into 54 genera.
The typical warblers are small birds belonging to the genus Sylvia in the "Old World warbler" family Sylviidae.
The parrotbills are a family, Paradoxornithidae, of passerine birds that are primarily native to East, Southeast and South Asia, with a single species in western North America, though feral populations exist elsewhere. They are generally small birds that inhabit reedbeds, forests and similar habitats. The traditional parrotbills feed mainly on seeds, e.g. of grasses, to which their robust bill, as the name implies, is well-adapted. Members of the family are usually non-migratory.
The white-eyes are a family, Zosteropidae, of small passerine birds native to tropical, subtropical and temperate Sub-Saharan Africa, southern and eastern Asia, and Australasia. White-eyes inhabit most tropical islands in the Indian Ocean, the western Pacific Ocean, and the Gulf of Guinea. Discounting some widespread members of the genus Zosterops, most species are endemic to single islands or archipelagos. The silvereye, Zosterops lateralis, naturally colonised New Zealand, where it is known as the "wax-eye" or tauhou ("stranger"), from 1855. The silvereye has also been introduced to the Society Islands in French Polynesia, while the Japanese white-eye has been introduced to Hawaii.
The Dartford warbler is a typical warbler from the warmer parts of western Europe and northwestern Africa. It is a small warbler with a long thin tail and a thin pointed bill. The adult male has grey-brown upperparts and is dull reddish-brown below except for the centre of the belly which has a dirty white patch. It has light speckles on the throat and a red eye-ring. The sexes are similar but the adult female is usually less grey above and paler below.
Pomatorhinus is a genus of scimitar babblers, jungle birds with long downcurved bills. These are birds of tropical Asia, with the greatest number of species occurring in hills of the Himalayas.
The wrentit is a small bird that lives in chaparral, oak woodlands, and bushland on the western coast of North America. It is the only species in the genus Chamaea.
The ashy-throated parrotbill is a parrotbill. In old sources, it may be called Alphonse's crow-tit; though superficially resembling a tit it is not a member of the Paridae. The native range of this species extends from south-west China to northern Vietnam, and it might have become naturalised in one area in Italy.
Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri. While more recent research suggests that its sister parvorder, Corvida, is not a monophyletic grouping, the Passerida as a distinct clade are widely accepted.
Turdoides is a genus of passerine birds in the laughingthrush family Leiothrichidae. The species are distributed across Africa and southern Asia and are typically fairly large, long-tailed birds which forage in noisy groups. The majority of species have drab brown or grey-brown plumage. Several species that were included in Turdoides in the past have been reassigned to Argya following a 2018 study that found multiple clades.
The bare-headed laughingthrush is a species of bird in the Old World babbler family Timaliidae. It is endemic to highland forests at elevations of 750–1,800 m (2,460–5,910 ft) in the mountain ranges of north-central Borneo, along with some outlying peaks. It is 25–26 cm (9.8–10.2 in) long, with both sexes similar in appearance. The head is brownish to greenish yellow and featherless. The area along the lower mandible has a bluish tinge. The rest of the body is dull blackish-brown tinged with grey. Juveniles have more feathers on the head, extending from the forehead to the crown.
The slender-billed scimitar babbler is a passerine bird in the Old World babbler family. It is found from the Himalayas to north-western Vietnam. Its natural habitat is subtropical or tropical moist montane forests.
Sylvioidea is a superfamily of passerine birds, one of at least three major clades within the Passerida along with the Muscicapoidea and Passeroidea. It contains about 1300 species including the Old World warblers, Old World babblers, swallows, larks and bulbuls. Members of the clade are found worldwide, but fewer species are present in the Americas.
Alcippe is a genus of passerine birds in the monotypic family Alcippeidae. The genus once included many other fulvettas and was previously placed in families Pellorneidae or Timaliidae.
The jungle babblers are a family, Pellorneidae, of mostly Old World passerine birds belonging to the superfamily Sylvioidea. They are quite diverse in size and coloration, and usually characterised by soft, fluffy plumage and a tail on average the length of their body, or longer. These birds are found in tropical zones, with the greatest biodiversity in Southeast Asia and the Indian subcontinent.
The laughingthrushes are a family, Leiothrichidae, of Old World passerine birds. The family contains 133 species which are divided into 16 genera. They are diverse in size and coloration. These are birds of tropical areas, with the greatest variety in Southeast Asia and the Indian subcontinent. The entire family used to be included in the Old World babbler family Timaliidae.