Palmichnium Temporal range: | |
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Trackway of P. palmatum at the Naturmuseum Senckenberg | |
Trace fossil classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Order: | † Eurypterida |
Ichnogenus: | † Palmichnium Richter, 1954 |
Type ichnospecies | |
†Palmichnium palmatum Richter, 1954 |
Palmichnium ("palm trace") is an ichnofossil genus, interpreted as a eurypterid trackway. It has been found by many places around the world, such as Australia, Canada, United States and Wales.
Its trackways consist of three or four subcircular tracks that are symmetrical around a midline impression that is arranged en echelon with a high angle to the midline. [1] [2]
The ichnogenus contains eight ichnospecies.
In 1975, Ann M. Anderson described a new ichnospecies of Petalichnus from the Table Mountain Sandstone of South Africa, P. capensis, Ordovician in time. Its ichnospecific name capensis refers to the type locality in the Cape Province of South Africa. Anderson noted that P. capensis was larger than the rest of the Petalichnus ichnospecies, as well as the presence of an unusual median drag line for the ichnogenus (although this was not present in all the specimens). [3] However, in 1999, Simon J. Braddy and John E. Almond formally reclassified some specimens of Petalichnus capensis to Palmichnium due to the possession of four tracks and a medium line, the rest of the specimens were referred to Petalichnus brandenburgensis. The trackway of P. capensis is defined as medium-sized (largest track 13.6 cm, 5.4 in wide) and consists of several symmetrical series of four tracks, each formed by a simple oval or tear-shaped mark with small impressions on the sides, sometimes bilobed (divided into two lobes) and intermittent formed by closely spaced series. These tracks were located at a low angle with respect to the sometimes present median line, which is believed to be due to the telson ("tail") touching the substrate. The bilobed marks is thought to be "eight"-shaped due to the long spines of the sixth appendage of an onychopterellid eurypterid, presumably Onychopterella augusti , using a swimming stroke-like motion. [4]
In 1983, Derek E. Gilmor Briggs and William D. Ian Rolfe described two new ichnospecies from Pennsylvania, P. kosinskiorum and P. stoermeri. The trace fossil of both ichnospecies were collected at a party from the Carnegie Museums of Pittsburgh in 1948. Its large size attracted the attention of the press, which attributed it to three-clawed amphibian hopping bipedally. The median line, the nature of the footprints and the lack of a suitable hopping vertebrate at the time rule out any hypothesis of an amphibian or vertebrate as the author of the footprints. In the case of invertebrates, it was concluded that only a large eurypterid that walked on six legs (hexapodous) could have produced the tracks. [5]
Braddy erected in 1995 P. macdonaldi for Lower Permian trackways found in the Hueco Formation of New Mexico, becoming the youngest species of the ichnogenus. The specimens, including holotype P23404, are housed in the New Mexico Museum of Natural History and Science. The ichnospecies is named in honor of Jerry MacDonald, the discoverer and local collector of the ichnofauna of the Robledo Mountains. The prints had a width of 25 mm (1 in) and consisted of symmetrical series of three to four pairs of impressions, large paddle-shaped outer tracks of 12 mm (0.5 in) in length and two or three internal linear impressions. In front of some outer tracks there is a slight curvilinear impression slightly convex outwards, possibly representing the recovery of a stroke of an appendage. There are also present inner tracks, small and linear scritches. An intermittent median groove preserved in the middle of the trackway is situated between the outer tracks. The producer of the footprints was a small arthropod with long and paddle-shaped posterior appendages and with three pairs of appendages in front, using an in-phase gait. Due to these characteristics it has been suggested that the producer was Adelophthalmus luceroensis , which has been found in nearby deposits (Madera Formation, central New Mexico). [6]
In 1843, the English geologist and paleontologist William Buckland described the now called BGS GSM 26037 (housed at the British Geological Survey in Nottingham, England) specimen as the result of the repeated impressions of three bony processes of the pectoral fin of an ambulatory (related or adapted to walk) fish, Icthyopatolites. The specimen was collected in Mostyn, Wales, and is Westphalian (Late Carboniferous) in age. However, in 1996, Braddy and Lyall I. Anderson recognized the track as clearly that of a moderately large (around 20 cm, 7.9 in long) hexapodous arthropod similar to the eurypterid traits that were described before in other papers, and named it Palmichnium pottsae. Although the leg morphology of the Carboniferous eurypterids is poorly known or completely unknown, the trackway fits with the maximum leg span of Adelophthalmus, approximately 10 cm (3.9 in). P. pottsae produced large trackways with asymmetric rows of three paired impressions along several irregular prints. It lacked a median line, but it has been suggested that this is due to the buoyancy of the water surrounding the abdomen of the animal or the active held up of the telson on the substrate without touching it, allowing better locomotion. The total length of the trackway was 37 cm (14.6 in). The outer tracks of the right and left series of the trackway were curvilinear or linear grooves, as well as the intermediate tracks, which were smaller than the outer ones. The internal tracks of both series were the smallest and with a bifid (cleft-like) form. There were also several irregular tracks that did not belong to any of the series. Although the assignment of P. pottsae to Palmichnium should mean the synonymy of Icthyopatolites with the first ichnogenus, Braddy and Anderson avoided formally synonymizing them because Buckland informally described Icthyopatolites and without an established diagnosis. [7] Early Devonian trackways have also been found in Alken an der Mosel, Germany. [8]
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event 251.9 million years ago.
Stylonurina is one of two suborders of eurypterids, a group of extinct arthropods commonly known as "sea scorpions". Members of the suborder are collectively and informally known as "stylonurine eurypterids" or "stylonurines". They are known from deposits primarily in Europe and North America, but also in Siberia.
Hibbertopterus is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of Hibbertopterus have been discovered in deposits ranging from the Devonian period in Belgium, Scotland and the United States to the Carboniferous period in Scotland, Ireland, the Czech Republic and South Africa. The type species, H. scouleri, was first named as a species of the significantly different Eurypterus by Samuel Hibbert in 1836. The generic name Hibbertopterus, coined more than a century later, combines his name and the Greek word πτερόν (pteron) meaning "wing".
Arthropleura is a genus of extinct millipede arthropods that lived in what is now North America and Europe around 345 to 290 million years ago, from the Viséan stage of the lower Carboniferous Period to the Sakmarian stage of the lower Permian Period. The species of the genus are the largest known land invertebrates of all time, and would have had few, if any, predators.
Chasmataspidids, sometime referred to as chasmataspids, are a group of extinct chelicerate arthropods that form the order Chasmataspidida. Chasmataspidids are probably related to horseshoe crabs (Xiphosura) and/or sea scorpions (Eurypterida), with more recent studies suggest that they form a clade (Dekatriata) with Eurypterida and Arachnida. Chasmataspidids are known sporadically in the fossil record through to the mid-Devonian, with possible evidence suggesting that they were also present during the late Cambrian. Chasmataspidids are most easily recognised by having an opisthosoma divided into a wide forepart (preabdomen) and a narrow hind part (postabdomen) each comprising 4 and 9 segments respectively. There is some debate about whether they form a natural group.
Diplichnites are arthropod trackways with two parallel rows of blunt to elongate, closely spaced tracks oriented approximately perpendicularly to the mid-line of the trackway. The term is more often used for the ichnofossils of this description; however, similar trackways from recent arthropods are sometimes given this name as well.
Campylocephalus is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of Campylocephalus have been discovered in deposits ranging from the Carboniferous period in the Czech Republic to the Permian period of Russia. The generic name is composed of the Greek words καμπύλος (kampýlos), meaning "curved", and κεφαλή (kephalē), meaning "head".
Onychopterella is a genus of predatory eurypterid, an extinct group of aquatic arthropods. Fossils of Onychopterella have been discovered in deposits from the Late Ordovician to the Late Silurian. The genus contains three species: O. kokomoensis, the type species, from the Early Pridoli epoch of Indiana; O. pumilus, from the Early Llandovery epoch of Illinois, both from the United States; and O. augusti, from the Late Hirnantian to Early Rhuddanian stages of South Africa.
Adelophthalmus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Adelophthalmus have been discovered in deposits ranging in age from the Early Devonian to the Early Permian, which makes it the longest lived of all known eurypterid genera, with a total temporal range of over 120 million years. Adelopththalmus was the final genus of the Eurypterina suborder of eurypterids and consisted the only known genus of swimming eurypterids from the Middle Devonian until its extinction during the Permian, after which the few surviving eurypterids were all walking forms of the suborder Stylonurina.
Euthycarcinoidea are an enigmatic group of extinct possibly amphibious arthropods that ranged from Cambrian to Triassic times. Fossils are known from Europe, North America, Argentina, Australia and Antarctica.
Mycteroptidae are a family of eurypterids, a group of extinct chelicerate arthropods commonly known as "sea scorpions". The family is one of three families contained in the superfamily Mycteropoidea, which in turn is one of four superfamilies classified as part of the suborder Stylonurina.
Pentecopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils have been registered from the Darriwilian age of the Middle Ordovician period, as early as 467.3 million years ago. The genus contains only one species, P. decorahensis, that is the oldest known eurypterid, surpassing other Ordovician eurypterids, such as Brachyopterus, in age by almost 9 million years. The generic name derives from the penteconter, a warship from ancient Greece, and the suffix -pterus, which means "wing" and is often used in other genus of eurypterids. The specific name refers to Decorah, Iowa, where Pentecopterus was discovered.
Adelophthalmidae is a family of eurypterids, an extinct group of aquatic arthropods. Adelophthalmidae is the only family classified as part of the superfamily Adelophthalmoidea, which in turn is classified within the infraorder Diploperculata in the suborder Eurypterina.
Eurypterina is one of two suborders of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". Eurypterine eurypterids are sometimes informally known as "swimming eurypterids". They are known from fossil deposits worldwide, though primarily in North America and Europe.
Wiedopterus is a genus of eurypterid, an extinct group of aquatic arthropods. The type and only species of Wiedopterus, W. noctua, is known from deposits of Early Devonian age in Germany. The generic name derives from the Wied river, which runs near the site of the initial discovery, and the species name, noctua, derives from Latin noctua (owl) which refers to the superficial resemblance of the carapace to an owl.
Ciurcopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Ciurcopterus have been discovered in deposits of Late Silurian age in North America. Classified as part of the family Pterygotidae, the genus contains two species, C. sarlei from Pittsford, New York and C. ventricosus from Kokomo, Indiana. The genus is named in honor of Samuel J. Ciurca, Jr., who has contributed significantly to eurypterid research by discovering a large amount of eurypterid specimens, including the four specimens used to describe Ciurcopterus itself.
Borchgrevinkium is an extinct genus of chelicerate arthropod. A fossil of the single and type species, B. taimyrensis, has been discovered in deposits of the Early Devonian period in the Krasnoyarsk Krai, Siberia, Russia. The name of the genus honors Carsten Borchgrevink, an Anglo-Norwegian explorer who participated in many expeditions to Antarctica. Borchgrevinkium represents a poorly known genus whose affinities are uncertain.
This timeline of eurypterid research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of eurypterids, a group of extinct aquatic arthropods closely related to modern arachnids and horseshoe crabs that lived during the Paleozoic Era.
Dromopus is a reptilian ichnogenus commonly found in assemblages of ichnofossils dating to the late Pennsylvanian to the late Permian. It has been found throughout Europe, as well as in the United States, Canada, and Morocco. Several ichnospecies have been named; only the type ichnospecies D. lacertoides is definitively recognized.
Tonganoxichnus is a Pennsylvanian to Permian trace fossil that has been found in North America.