Parenchymia

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Parenchymia
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Clade: Bilateria
Clade: Nephrozoa
(unranked): Protostomia
(unranked): Spiralia
Superphylum: Lophotrochozoa
Clade: Parenchymia
Phyla

Parenchymia is a proposed clade encompassing Nemertea and Platyhelminthes. The clade was suggested in a 2019 study showing the clade as a sister to Annelida, with both groups together being sister to Lophophorata. The synapomorphies include similar organization of ciliary bands in the larva of both groups, as well as a lack of chitin, and is named for the abundant parenchyma found between the body wall and internal organs of both. [1] A similar taxon, with Mollusca being likewise linked together with Symbion and Entoprocta, has been proposed in a clade called Tetraneuralia. [2]

Phylogeny

  Protostomia  

Ecdysozoa Aptostichus simus Monterey County.jpg

  Spiralia   (s.l.)  /

Gnathifera Pseudosagitta maxima 31349361.png



  Lophotrochozoa  / 
  Tetraneuralia  
 

Mollusca Grapevinesnail 01a.jpg

Entoprocta Barentsia laxa 1498941 (no background).png


 
 

Gastrotricha Diuronotus aspetos 1.png

  Lophophorata  
 
 

Ectoprocta Bugulina flabellata 272067784.png

Phoronida Phoronopsis harmeri IZ 1643662.png

Brachiopoda LingulaanatinaAA (cropped).JPG

Annelida Polychaeta (no) 2.jpg

  Parenchymia  
 

Platyhelminthes Pseudobiceros bedfordi 13376124.png

Nemertea Micrura verrilli 218962819.png

  Platytrochozoa  / 
Spiralia   (s.s.)
  Gnathospiralia  

Related Research Articles

<span class="mw-page-title-main">Ecdysozoa</span> Superphylum of protostomes including arthropods, nematodes and others

Ecdysozoa is a group of protostome animals, including Arthropoda, Nematoda, and several smaller phyla. The grouping of these animal phyla into a single clade was first proposed by Eernisse et al. (1992) based on a phylogenetic analysis of 141 morphological characters of ultrastructural and embryological phenotypes. This clade, that is, a group consisting of a common ancestor and all its descendants, was formally named by Aguinaldo et al. in 1997, based mainly on phylogenetic trees constructed using 18S ribosomal RNA genes.

<span class="mw-page-title-main">Trochozoa</span> Taxonomic clade

The Trochozoa are a proposed Lophotrochozoa clade that is a sister clade of Bryozoa. The clade would include animals in five phyla: the Nemertea, the Annelida, the Mollusca, and the two Brachiozoan phyla, Brachiopoda and Phoronida.

<span class="mw-page-title-main">Ornithischia</span> Extinct clade of dinosaurs

Ornithischia is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name Ornithischia, or "bird-hipped", reflects this similarity and is derived from the Greek stem ornith- (ὀρνιθ-), meaning "bird", and ischion (ἴσχιον), meaning "hip". However, birds are only distantly related to this group as birds are theropod dinosaurs. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs, the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous pelts, and there is much debate over whether these filaments found in specimens of Tianyulong, Psittacosaurus, and Kulindadromeus may have been primitive feathers.

<span class="mw-page-title-main">Lophotrochozoa</span> Superphylum of animals

Lophotrochozoa is a clade of protostome animals within the Spiralia. The taxon was established as a monophyletic group based on molecular evidence. The clade includes animals like annelids, molluscs, bryozoans, brachiopods, and platyhelminthes.

<span class="mw-page-title-main">Paulowniaceae</span> Family of trees

Paulowniaceae are a family of flowering plants within the Lamiales. They are a monophyletic and monogeneric family of trees with currently 7 confirmed species. They were formerly placed within Scrophulariaceae sensu lato, or as a segregate of the Bignoniaceae.

<span class="mw-page-title-main">Asterids</span> Clade of eudicot angiosperms

In the APG IV system (2016) for the classification of flowering plants, the name asterids denotes a clade. Asterids is the largest group of flowering plants, with more than 80,000 species, about a third of the total flowering plant species. Well-known plants in this clade include the common daisy, forget-me-nots, nightshades, the common sunflower, petunias, yacon, morning glory, lettuce, sweet potato, coffee, lavender, lilac, olive, jasmine, honeysuckle, ash tree, teak, snapdragon, sesame, psyllium, garden sage, table herbs such as mint, basil, and rosemary, and rainforest trees such as Brazil nut.

<span class="mw-page-title-main">Drepanosaur</span> Extinct clade of reptiles

Drepanosaurs are a group of extinct reptiles that lived between the Carnian and Rhaetian stages of the late Triassic Period, approximately between 230 and 210 million years ago. The various species of drepanosaurid were characterized by specialized grasping limbs and often prehensile tails, adaptions for arboreal (tree-dwelling) and fossorial (digging) lifestyles, with some having also been suggested to be aquatic. Fossils of drepanosaurs have been found in Arizona, New Mexico, New Jersey, Utah, England, and northern Italy. The name is taken from the family's namesake genus Drepanosaurus, which means "sickle lizard," a reference to their strongly curved claws.

<span class="mw-page-title-main">Deuterostome</span> Superphylum of bilateral animals

Deuterostomes are bilaterian animals of the superphylum Deuterostomia, typically characterized by their anus forming before the mouth during embryonic development. Deuterostomia is further divided into 4 phyla; Chordata, Echinodermata, Hemichordata, the extinct Vetulicolia known from Cambrian fossils. The extinct clade Cambroernida is also though to be a member of Deuterostomia.

<span class="mw-page-title-main">SAR supergroup</span> Eukaryotes superphylum

SAR or Harosa is a highly diverse clade of eukaryotes, often considered a supergroup, that includes stramenopiles (heterokonts), alveolates, and rhizarians. It is a node-based taxon, including all descendants of the three groups' last common ancestor, and comprises most of the now-rejected Chromalveolata. Their sister group has been found to be telonemids, with which they make up the TSAR clade.

<span class="mw-page-title-main">Gnathifera (clade)</span> Taxonomic clade

Gnathifera is a clade of generally small spiralians characterized by complex jaws made of chitin. It comprises the phyla Gnathostomulida, Rotifera and Micrognathozoa. Chaetognatha has recently been recognised as closely related to the group, with it either being included within Gnathifera or the broader group Chaetognathifera. It may also include the Cycliophora.

<span class="mw-page-title-main">Spiralia</span> Clade of protosomes with spiral cleavage during early development

The Spiralia are a morphologically diverse clade of protostome animals, including within their number the molluscs, annelids, platyhelminths and other taxa. The term Spiralia is applied to those phyla that exhibit canonical spiral cleavage, a pattern of early development found in most members of the Lophotrochozoa.

<span class="mw-page-title-main">Aetosaurinae</span> Extinct subfamily of reptiles

Aetosaurinae is one of the two main clades of aetosaurs, the other being Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Aetosaurus than Desmatosuchus. Aetosaurinae currently comprises Aetosaurus, similar forms such as Coahomasuchus and Stenomyti, and the widespread and successful aetosaur clade Typothoracinae.

<span class="mw-page-title-main">Nephrozoa</span> Clade of animals

Nephrozoa is a proposed major clade of bilaterian animals. It includes all bilaterians other than Xenacoelomorpha. It contrasts with the Xenambulacraria hypothesis, which instead posits that Xenocoelomorpha is nested within Bilateria as most closely related to Ambulacraria. Which hypothesis is correct is controversial. Authors supporting the Xenambulacraria hypothesis have suggested that the genetic evidence used to support Nephrozoa may be due to systematic error.

<span class="mw-page-title-main">Scincogekkonomorpha</span> Clade of lizards

Scincogekkonomorpha is a proposed clade of lizards that includes scleroglossans and all lizards more closely related to scleroglossans than to iguanians. These "stem" scleroglossans include extinct lizards from the Late Jurassic and Early Cretaceous such as Bavarisaurus, Eichstaettisaurus, Liushusaurus, and Scandensia. Scincogekkonomorpha was named in 1961 and is now occasionally used as a stem-based taxon in contrast to the node-based taxon Scleroglossa. According to phylogenies based on morphological characteristics, Scincogekkonomorpha is the sister taxon of Iguania and together they make up crown group Squamata, the smallest clade including all living snakes and lizards. The grouping has not been recovered as monophyletic in recent molecular studies, with Iguania generally found deeply nested within Squamata.

<span class="mw-page-title-main">Ichthyosauromorpha</span> Clade of marine reptiles

The Ichthyosauromorpha are an extinct clade of marine reptiles consisting of the Ichthyosauriformes and the Hupehsuchia, living during the Mesozoic.

<span class="mw-page-title-main">Pantestudines</span> Clade of reptiles

Pantestudines or Pan-Testudines is the group of all reptiles more closely related to turtles than to any other living animal. It includes both modern turtles and all of their extinct relatives. Pantestudines with a complete shell are placed in the clade Testudinata.

The Platytrochozoa are a proposed basal clade of spiralian animals as the sister group of the Gnathifera. The Platytrochozoa were divided into the Rouphozoa and the Lophotrochozoa. A more recent study suggests that the mesozoans also belong to this group of animals, as sister of the Rouphozoa.

<span class="mw-page-title-main">Xenambulacraria</span> Animal clade containing xenoturbellids, acoelomorphs, echinoderms and hemichordates

Xenambulacraria is a proposed clade of animals with bilateral symmetry as an embryo, consisting of the Xenacoelomorpha and the Ambulacraria.

<span class="mw-page-title-main">Colossosauria</span> Extinct clade of dinosaurs

Colossosauria is a clade of titanosaur sauropods from the latest Early Cretaceous through the Late Cretaceous of South America. The group was originally named by Bernardo González-Riga et al. in 2019 and defined as the "most inclusive clade containing Mendozasaurus neguyelap but not Saltasaurus loricatus or Epachthosaurus sciuttoi". The clade contains different taxa depending on the phylogenetic analysis used, in the defining paper the only subgroups were Rinconsauria and Lognkosauria, but alternate phylogenies published previously had also included various similar titanosaurs such as Aeolosaurus, Bonitasaura, Drusilasaura, Overosaurus and Quetecsaurus. The phylogenetic analysis of González-Riga et al. (2019) placed Colossosauria as sister taxa to Epachthosaurus, Pitekunsaurus and a larger clade including Saltasauridae.

<span class="mw-page-title-main">Centroneuralia</span> Animal clade containing chordates and protostomes

Centroneuralia is a proposed clade of animals with bilateral symmetry as an embryo, consisting of the Chordata and Protostomia, united by the presence of a central nervous system. An alternative to the traditional protostome-deuterostome dichotomy, it has found weak support in several studies. Under this hypothesis, Centroneuralia would be sister to Xenambulacraria at the base of Bilateria.

References

  1. Marlétaz, Ferdinand; Peijnenburg, Katja T.C.A.; Goto, Taichiro; Satoh, Noriyuki; Rokhsar, Daniel S. (2019-01-21). "A New Spiralian Phylogeny Places the Enigmatic Arrow Worms among Gnathiferans" (PDF). Current Biology. 29 (2): 312–318.e3. doi: 10.1016/j.cub.2018.11.042 . ISSN   0960-9822. PMID   30639106. S2CID   58562919.
  2. Bleidorn, Christoph (2019-08-22). "Recent progress in reconstructing lophotrochozoan (spiralian) phylogeny". Organisms Diversity & Evolution. Springer Science and Business Media LLC. 19 (4): 557–566. doi:10.1007/s13127-019-00412-4. ISSN   1439-6092. S2CID   201283619.