Pinnipedimorphs | |
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Artist reconstruction of Enaliarctos emlongi | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Clade: | Pan-Pinnipedia |
Clade: | Pinnipedimorpha Berta et al., 1989 |
Subtaxa | |
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Pinnipedimorpha is a clade of arctoid carnivorans that is defined to include the last common ancestor of Phoca and Enaliarctos , and all descendants of that ancestor. [1] Scientists still debate on which lineage of arctoid carnivorans are the closest relatives to the pinnipedimorphs, being more closely related to musteloids. [2] [3] [4] [5] [6]
Below is an overall phylogeny of the taxa covered in the article followed after a composite tree in Berta et al. (2018) [1] and a total-evidence (combined molecular-morphological) dataset in Paterson et al. (2020): [7]
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Carnivora is an order of placental mammals that have specialized in primarily eating flesh, whose members are formally referred to as carnivorans. The order Carnivora is the fifth largest order of mammals, comprising at least 279 species.
The earless seals, phocids, or true seals are one of the three main groups of mammals within the seal lineage, Pinnipedia. All true seals are members of the family Phocidae. They are sometimes called crawling seals to distinguish them from the fur seals and sea lions of the family Otariidae. Seals live in the oceans of both hemispheres and, with the exception of the more tropical monk seals, are mostly confined to polar, subpolar, and temperate climates. The Baikal seal is the only species of exclusively freshwater seal.
An eared seal, otariid, or otary is any member of the marine mammal family Otariidae, one of three groupings of pinnipeds. They comprise 15 extant species in seven genera and are commonly known either as sea lions or fur seals, distinct from true seals (phocids) and the walrus (odobenids). Otariids are adapted to a semiaquatic lifestyle, feeding and migrating in the water, but breeding and resting on land or ice. They reside in subpolar, temperate, and equatorial waters throughout the Pacific and Southern Oceans, the southern Indian, and Atlantic Oceans. They are conspicuously absent in the north Atlantic.
Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs, bears, raccoons, and mustelids. The Pinnipedia are also assigned to this group. The center of diversification for the Caniformia is North America and northern Eurasia. Caniformia stands in contrast to the other suborder of Carnivora, the Feliformia, the center of diversification of which was in Africa and southern Asia.
Ailuridae is a family in the mammal order Carnivora. The family consists of the red panda and its extinct relatives.
Ferae is a mirorder of placental mammals from grandorder Ferungulata, that groups together clades Pan-Carnivora, which includes modern carnivorans, and Pholidotamorpha, which includes pangolins.
Eupleridae is a family of carnivorans endemic to Madagascar and comprising 10 known living species in seven genera, commonly known as euplerids, Malagasy mongooses or Malagasy carnivorans. The best known species is the fossa, in the subfamily Euplerinae. All species of Euplerinae were formerly classified as viverrids, while all species in the subfamily Galidiinae were classified as herpestids.
Galidiinae is a subfamily of carnivorans that is restricted to Madagascar and includes six species classified into four genera. Together with the three other species of indigenous Malagasy carnivorans, including the fossa, they are currently classified in the family Eupleridae within the suborder Feliformia. Galidiinae are the smallest of the Malagasy carnivorans, generally weighing about 600 to 900 g. They are agile, short-legged animals with long, bushy ringed tails.
Potamotherium an extinct genus of caniform carnivoran from the Miocene epoch of France and Germany. It has historically been assigned to the family Mustelidae, but more recent studies suggest that it represents a primitive relative of pinnipeds
Enaliarctos is an extinct genus of pinnipedimorph, and may represent the ancestor to all pinnipeds. Prior to the discovery of Puijila, the five species in the genus Enaliarctos represented the oldest known pinnipedimorph fossils, having been recovered from late Oligocene and early Miocene strata of California and Oregon.
Carnivoramorpha is a clade of placental mammals of clade Pan-Carnivora from mirorder Ferae, that includes the modern order Carnivora and its extinct stem-relatives.
Ursoidea is a superfamily of arctoid carnivoran mammals that includes the families Subparictidae, Amphicynodontidae, and Ursidae which the last family includes the extant lineages of bears as well the extinct Hemicyoninae and Ursavinae. The interrelationships of ursoids has had slight arrangements. In the past it was thought the extinct Amphicyonidae were stem-bears based on morphological analysis of the ear region, though the most recent publications on early amphicyonids suggests they were basal caniforms. The amphicynodontids are sometimes classified as either a subfamily of bears, a paraphyletic assemblage of early bears, or even stem-pinnipeds. The subparictids were previously classified as amphicynodontine/ids. The hemicyonines have been occasionally reclassified as a separate family.
Musteloidea is a superfamily of carnivoran mammals united by shared characteristics of the skull and teeth. Musteloids are the sister group of pinnipeds, the group which includes seals.
Arctoidea is a clade of mostly carnivorous mammals which include the extinct Hemicyonidae (dog-bears), and the extant Musteloidea, Pinnipedia, and Ursidae (bears), found in all continents from the Eocene, 46 million years ago, to the present. The oldest group of the clade is the bears, as their CMAH gene is still intact. The gene became non-functional in the common ancestor of the Mustelida. Arctoids are caniforms, along with dogs (canids) and extinct bear dogs (Amphicyonidae). The earliest caniforms were superficially similar to martens, which are tree-dwelling mustelids. Together with feliforms, caniforms compose the order Carnivora; sometimes Arctoidea can be considered a separate suborder from Caniformia and a sister taxon to Feliformia.
The Astoria Formation is a geologic formation in Washington state & Oregon. It preserves fossils dating back to the early to middle Miocene.
Amphicynodontidae is a probable clade of extinct arctoids. While some researchers consider this group to be an extinct subfamily of bears, a variety of morphological evidence links amphicynodontines with pinnipeds, as the group were semi-aquatic otter-like mammals. In addition to the support of the pinniped–amphicynodontine clade, other morphological and some molecular analyses support bears being the closest living relatives to pinnipeds. According to McKenna and Bell (1997) Amphicynodontinae are classified as stem-pinnipeds in the superfamily Phocoidea. Fossils of these mammals have been found in Europe, North America and Asia. Amphicynodontines should not be confused with Amphicyonids (bear-dogs), a separate family of Carnivora which is a sister clade to arctoids within the caniforms, but which may be listed as a clade of extinct arctoids in older publications.
Annalisa Berta is an American paleontologist and professor emerita in the Department of Biology at San Diego State University.
Otarioidea is a superfamily of pinnipeds that includes the families Odobenidae, Otariidae and their stem-relatives. In the past when the pinnipeds were considered to be a diphyletic group of marine mammals, a few points of cranial and dental morphology suggested that the otarioids originated from a line of bears. One extinct family, Enaliarctidae, was postulated to be otarioids that were a transitional clade between Hemicyoninae and Otariidae. Recent comprehensive studies have, however, since the 1990s found pinnipeds to be a monophyletic clade of aquatic arctoids. There are a few authorities that place desmatophocids and odobenids as sister taxa to Phocidae in the clade Phocomorpha based on a few minor physiological features.
Subparictidae is an extinct family of early Paleogene arctoid carnivorans endemic to North America that closely related to bears. They were small, raccoon-like mammals that lived from the Eocene to the early Miocene. This family includes a handful of genera such as Subparictis, Parictis, Nothocyon, and Eoarctos.