Praenuculidae

Last updated

Praenuculidae
Temporal range: Ordovician - Devonian 487–391  Ma
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Bivalvia
Order: Nuculida
Superfamily: Nuculoidea
Family: Praenuculidae
McAlester, 1969
Subfamilies

See text

Praenuculidae is an extinct family of prehistoric bivalves in the superfamily Nuculoidea. [1] Praenuculidae species lived from the early Ordovician, Arenig stage through the Early Devonian Emsian stage. [2] Praenuculidae fossils are found worldwide, present on every continent except Antarctica. [2] Species in this family are thought to have been sessile, attached to the substrate in shallow infaunal marine water environments, where they formed shells of an aragonite composition. The family Praenuculidae was named by A. Lee McAlester in 1969. [2]

Contents

Description

Praenuculidae first emerged in the early Ordovician and diversified from around 6 genera in the early Orodvician to a maximum of thirteen genera by the Late Ordovician. [3] As a result of the Ordovician–Silurian extinction event the family was reduced to three genera during the Silurian and by the end of the Devonian the family was entirely extinct. [3]

The family is composed of up to seventeen genera, most divided between the two described subfamilies erected by Teresa M. Sánchez in 1999. [4] The structure of the chevroned hinge teeth is the dominant feature by which members of Praenuculidae are divided between the two subfamilies. The majority of Praenuculidae genera possess teeth with a chevron concavity that faces towards the outer sides of the shell and a chevron point facing the umbo. [4] These genera are grouped into the larger subfamily, Praenuculinae. Three of the genera are placed into the second subfamily, Concavodontinae, based on the teeth having chevron concavities which face in the reverse, towards the center of the hinge and points facing the outside edges of the shell. [4]

In the original 1999 description of the subfamilies, Cardiolaria was not placed into a specific subfamily due to the uncertain family affinities of the genus. Cardiolaria was placed in Praenuculidae by McAlester in 1969 with the original description of the family. It was suggested by John C. Cope in 1997 that the genus may belong elsewhere in the subclass Protobranchia, [1] formerly called Palaeotaxodonta. Both Deceptrix and Cardiolaria are currently accepted as members of the family Cardiolariidae erected by Cope in 1997. [5] [1] The genus Eritropis was formerly included in the family, however it was moved to a new family, Eritropidae by John C. Cope in 2000. [4] [6]

Genera

A number of genera which were placed in the family have been moved to other families. They include:

Related Research Articles

<span class="mw-page-title-main">Spiriferida</span> Order of brachiopods

Spiriferida is an order of extinct articulate brachiopod fossils which are known for their long hinge-line, which is often the widest part of the shell. In some genera it is greatly elongated, giving them a wing-like appearance. They often have a deep fold down the center of the shell. The feature that gives the spiriferids their name ("spiral-bearers") is the internal support for the lophophore; this brachidium, which is often preserved in fossils, is a thin ribbon of calcite that is typically coiled tightly within the shell.

<span class="mw-page-title-main">Strophomenida</span> Extinct order of brachiopods

Strophomenida is an extinct order of articulate brachiopods which lived from the lower Ordovician period to the mid Carboniferous period. Strophomenida is part of the extinct class Strophomenata, and was the largest known order of brachiopods, encompassing over 400 genera. Some of the largest and heaviest known brachiopod species belong to this class. Strophomenids were among the most diverse and abundant brachiopods during the Ordovician, but their diversity was strongly impacted at the Late Ordovician mass extinction. Survivors rediversified into new morphologies in the Silurian, only to be impacted once again at the Late Devonian mass extinction. However, they still survived till the mid Carboniferous.

<span class="mw-page-title-main">Rhynchonelliformea</span> Subphylum of brachiopods

Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.

Pojetaia is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early to Middle Cambrian fossils found in North America, Greenland, Europe, North Africa, Asia, and Australia. The genus currently contains two accepted species, Pojetaia runnegari, the type species, and Pojetaia sarhroensis, though up to seven species have been proposed. The genera Buluniella, Jellia, and Oryzoconcha are all considered synonyms of Pojetaia.

<span class="mw-page-title-main">Fordillidae</span> Extinct family of bivalves

Fordillidae is an extinct family of early bivalves and one of two families in the extinct superfamily Fordilloidea. The family is known from fossils of early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia. The family currently contains two genera, Fordilla and Pojetaia, each with up to three described species. Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish Small shelly fauna.

Concavodonta is an extinct genus of early bivalve in the extinct family Praenuculidae. The genus is one of three genera in the subfamily Concavodontinae. Concavodonta is known solely from late Ordovician, Caradoc epoch, fossils found in Europe and South America. The genus currently contains three accepted species, Concavodonta imbricata, Concavodonta ovalis and the type species Concavodonta ponderata.

Hemiconcavodonta is an extinct genus of bivalve in the extinct family Praenuculidae. The genus is one of three genera in the subfamily Concavodontinae. Hemiconcavodonta is known solely from late Ordovician, Caradoc epoch, fossils found in South America. The genus currently contains a single accepted species, Hemiconcavodonta minuta.

Concavodontinae is an extinct subfamily of prehistoric bivalves in the family Praenuculidae. Concavodontinae species lived from the middle Ordovician, Caradoc epoch through the late Ordovician Ashgill epoch. Concavodontinae fossils are found in Europe and South America, and species are thought to have been stationary attached to substrate in shallow infaunal marine water environments where they formed shells of an aragonite composition. The subfamily Concavodontinae was named by Teresa M. Sánchez in 1999.

Emiliodonta is an extinct genus of bivalve in the extinct family Praenuculidae. The genus is one of three genera in the subfamily Concavodontinae. Emiliodonta is known solely from late Ordovician, Caradocian epoch, fossils found in South America. The genus contains a single accepted species, Emiliodonta cuerdai.

Praenuculinae is an extinct subfamily of prehistoric bivalves in the family Praenuculidae. Praenuculinae species lived from the middle Ordovician through the late Devonian. Praenuculinae fossils are found in Europe, Africa, North America and South America, and species are thought to have been stationary attached to substrate in shallow infaunal marine water environments where they formed shells of an aragonite composition. The subfamily Praenuculinae was named by Teresa M. Sánchez in 1999.

Cuyopsis is an extinct genus of bivalve in the extinct family Praenuculidae. The genus is one of eleven genera in the subfamily Praenuculinae. It is one of three Praenuculinae genera known solely from late Ordivician, Caradoc epoch, fossils found in South America. Cuyopsis currently contains a single accepted species, Cuyopsis symmetricus.

Villicumia is an extinct genus of bivalve in the extinct family Praenuculidae. The genus is one of eleven genera in the subfamily Praenuculinae. It is one of three Praenuculinae genera known solely from late Ordovician, Caradoc epoch, fossils found in South America. Villicumia currently contains a single accepted species, Villicumia canteraensis.

Trigonoconcha is an extinct genus of bivalve in the extinct family Praenuculidae. The genus is one of eleven genera in the subfamily Praenuculinae. It is one of three Praenuculinae genera known solely from Late Ordivician, Caradoc epoch, fossils found in South America. Trigonoconcha currently contains a single accepted species, Trigonoconcha acuta.

<i>Similodonta</i> Extinct genus of bivalves

Similodonta is an extinct genus of early bivalve in the extinct family Praenuculidae. The genus is one of eleven genera in the subfamily Praenuculinae. Similodonta is known from Middle Ordovician through Middle Silurian fossils found in Europe and North America. The genus currently contains eight accepted species, Similodonta ceryx, Similodonta collina, Similodonta djupvikensis, Similodonta magna, Similodonta recurva, Similodonta spjeldnaesi, Similodonta wahli and the type species Similodonta similis.

<i>Panenka</i> (bivalve) Genus of bivalves

Panenka is a genus of fossil saltwater clams, marine bivalve molluscs in the family Praecardiidae. Like most bivalves, these molluscs were suspension feeders. They lived in the Devonian Period.

<span class="mw-page-title-main">Trimerellida</span> Extinct order of brachiopods

Trimerellida is an extinct order of craniate brachiopods, containing the sole superfamily Trimerelloidea and the families Adensuidae, Trimerellidae, and Ussuniidae. Trimerellidae was a widespread family of warm-water brachiopods ranging from the Middle Ordovician to the late Silurian (Ludlow). Adensuidae and Ussuniidae are monogeneric families restricted to the Ordovician of Kazakhstan. Most individuals were free-living, though some clustered into large congregations similar to modern oyster reefs.

Craniopsidae is an extinct family of craniiform brachiopods which lived from the mid-Cambrian to the Lower Carboniferous (Tournaisian). It is the only family in the monotypic superfamily Craniopsoidea and the monotypic order Craniopsida. If one includes the ambiguous Cambrian genus Discinopsis, craniopsids were the first craniiforms to appear, and may be ancestral to craniids and trimerellides. An even earlier Cambrian genus, Heliomedusa, has sometimes been identified as a craniopsid. More recently, Heliomedusa has been considered a stem-group brachiopod related to Mickwitzia.

<span class="mw-page-title-main">Adelophthalmidae</span> Family of eurypterids

Adelophthalmidae is a family of eurypterids, an extinct group of aquatic arthropods. Adelophthalmidae is the only family classified as part of the superfamily Adelophthalmoidea, which in turn is classified within the infraorder Diploperculata in the suborder Eurypterina.

<span class="mw-page-title-main">Orthotetida</span> Extinct order of marine lamp shells

The orthotetides (Orthotetida) are an extinct order of brachiopods in the class Strophomenata. Though not particularly diverse or abundant relative to strophomenides (Strophomenida) or productides (Productida), orthotetides were nevertheless the longest-lasting order of strophomenates, surviving from the Middle Ordovician (“Llanvirn”) up until the Late Permian. Externally, many orthotetides are difficult to distinguish from strophomenides. Most fundamental differences between the two orders are internal: orthotetides have more elaborate cardinal processes and a greater diversity of shell microstructure.

Askerina is an extinct genus of atrypid brachiopods from subfamily Atrypinae that lived in end-Ordovician (Hirnantian) and Silurian (Aeronian). The type and only known species is Askerina cymbula. Its fossils are known only from the lower parts of the Solvik Formation in central Oslo region, Norway.

References

  1. 1 2 3 Carter, J.G.; et al. (2011). "A Synoptical Classification of the Bivalvia (Mollusca)" (PDF). Paleontological Contributions. 4: 1–47.
  2. 1 2 3 The Paleobiology Database Praenuculidae entry accessed 11 January 2012
  3. 1 2 Nevesskaya, L.A. (2008). "Dynamics of Taxonomic Diversity of Bivalves in the Phanerozoic". Paleontological Journal. 42 (4): 335–342. doi:10.1134/S0031030108040011. S2CID   84204347.
  4. 1 2 3 4 Sánchez, T.M. (1999). "New Late Ordovician (Early Caradoc) Bivalves from the Sierra de Villicum (Argentine Precordillera)". Journal of Paleontology. 73 (1): 66–76. doi:10.1017/S0022336000027554. JSTOR   1306745. S2CID   133414164.
  5. The Paleobiology Database Cardiolariidae entry accessed 5 February 2012
  6. Zong-Jie, F.; Cope, J.C.W. (2004). "Early Ordovician bivalves from Dali, West Yunnan, China". Palaeontology. 47 (5): 1121–1158. doi:10.1111/j.0031-0239.2004.00403.x. S2CID   140648734.