Pseudohaloritidae

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Pseudohaloritidae
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Ammonoidea
Order: Goniatitida
Superfamily: Pseudohaloritoidea
Family: Pseudohaloritidae
Miller & Furnish, 1957
Subfamilies

Pseudohaloritidae is the larger of two families that form the goniatitid superfamily Pseudohaloritoidea, the other being the monogenerc Maximitidae. They are part of the vast array of shelled cephalopods known as ammonoids that are more closely related to squids, belemnites, octopuses, and cuttlefish, than to the superficially similar Nautilus . [1]

The Pseudohaloritide which now contains some 14 genera in three subfamilies is characterized by small, subdiscoidal to subglobular, involute shells, the surface of which may be smooth or with coarse longitudinal lirae and/or transverse ribs. The siphuncle is retrosiphonitic, a hold-over character from the nautiloids, usually subcentral or situated within dorsal septal flexure but ventral-marginal in first and second whorls. Sutures have four pairs of lobes, the ventral one being rounded and spatulate, the interior three clustered and narrow. External lobes are either smooth are variably serrate. [1] [2]

The Pseudohaloritidae was established by Miller and Furnish (1957) [2] for three related genera with similar sutures and aberrant siphuncles that are removed from the ventral margin. One, Maximites, has been put in a family of its own, the Maximitidae. The other two, Neoaganides and Pseudohalorites are retained in the Pseudohaloritidae, to which has been added about a dozen more genera, mostly from China. [3]

Maximites differs in having a bifid ventral lobe and a siphuncle that is ventral, but not marginal. It is also older, as early as the Middle Pennsylvanian (m U Carb) rather than from the Upper Pennsylvanian (uU Carb) when pseudohaloritids, as emended, make their first appearance. [4] Maximites is thought to have given rise to the Neoaganides and thus to the Pseudohaloritidae, and in turn may have been derived from Imitoceras. On the other hand, the suture indicates that Neoaganides may have developed directly from Imitoceras. The ventral lobe of Neoaganides is like that of Imitoceras, smooth and rounded, but the lateral lobes differ from those of both Imitoceras and Maximites in being pointed. [2]

The aberrant position of the siphuncle well away from the ventral margin and in some near the dorsum is characteristic of, although not unique to, this family. However no link is indicated between these Pennsylvanian and Permian forms and the Upper Devonian Clymeniids with their well established dorso-marginal siphuncles. [2]

Related Research Articles

Ammonoidea Extinct subclass of cephalopod molluscs

Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids than they are to shelled nautiloids such as the living Nautilus species. The earliest ammonites appear during the Devonian, and the last species vanished in the Cretaceous–Paleogene extinction event.

Goniatite order of molluscs (fossil)

Goniatids, informally goniatites, are ammonoid cephalopods that form the order Goniatitida, derived from the more primitive Agoniatitida during the Middle Devonian some 390 million years ago. Goniatites (goniatitids) survived the Late Devonian extinction to flourish during the Carboniferous and Permian only to become extinct at the end of the Permian some 139 million years later.

Yinoceras is a genus of middle Permian goniatitid ammonite, the type genus for the subfamily Yinoceratinae of the family Pseudohaloritidae.

Dimeroceratidae is one of three families in the Dimeroceratoidea, a goniatid superfamily included in the Ammonoidea; extinct shelled cephalopods with adorally convex septa and usually narrow ventro-marginal siphuncles.

Gastrioceratoidea is one of seventeen superfamilies in the suborder Goniatitina, ammonoid cephalopods from the Late Paleozoic.

Adrianitidae is a family in the Adrianitaceae, a superfamily of ammonites in the cephalopod order, Goniatitida, known from the Middle Pennsylvanian to the Middle Permian.

Thalassoceratidae a family of late Paleozoic ammonites included in the goniatitid superfamily Thalassoceratoidea along with the Bisatoceratidae. Some eight genera are included, although the specific number and exactly which depends on the particular classification.

Medlicottiidae is a family of ammonoid cephalopods belonging to the Prolecanitida, known from the Upper Carboniferous (Pennsylvanian) to the Early Triassic.

Prolecanitida is an order of extinct ammonoid cephalopods with discoidal to thinly lenticular shells with goniatitic or ceratitic sutures and which retained the simple retrochoanitic siphuncle with backward extending septal necks. As typical for ammonoids the siphuncle is along the ventral margin. Prolecanitids form a relatively small and stable order within the Ammonoidea with 43 named genera and about 1250 species, but with a long-ranging lineage of about 108 m.y. stretching from the Lower Carboniferous to the Triassic. Although not as diverse as their goniatitid contemporaries, the Prolecanatida provided the stock from which all later Mesozoic ammonoids were derived.

Permoceras, the sole member of the family Permoceratidae, is a genus of coiled nautiloids with a smooth, compressed involute shell, whorls higher than wide, earlier whorls hidden from view. The venter is rounded as are the ventral and umbilical shoulders, the flanks flattened. The siphuncle is ventrally subcentral. The suture, which is most characteristic, has a deep, narrow pointed ventral lobe and large, asymmetrical pointed lobes on either side.

The Clydonautiloidea are a superfamily within the nautiloid order Nautilida characterized by smooth, generally globular, shells with nearly straight sutures, in early forms, but developing highly differentiated sutures in some later forms. Where known, the siphuncle tends to be central to subcentral.

Trigonoceratoidea superfamily of molluscs

The Trigonoceratoidea are a superfamily within the Nautilida that ranged from the Devonian to the Triassic, thought to have contained the source for the Nautilaceae in which Nautilus is found.

Syringonautilidae is a family of Nautiloidea from the middle to late Triassic. Syringonautilidae comprise the last of the Trigonoceratoidea and are the source for the Nautilaceae which continued the Nautiloidea through the Mesozoic and into the Cenozoic right down to the recent. Syringonautilidae is a strictly Triassic family, derived early in the Triassic from the Grypoceratidae.

Grypoceratidae family of molluscs

Grypoceratidae is the longest-lived family of the Trigonoceratoidea, or of the near equivalent Centroceratina; members of the Nautilida from the Upper Paleozoic and Triassic.

The Centroceratidae is the ancestral family of the Trigonoceratoidea and of the equivalent Centroceratina; extinct shelled cephalopods belonging to the order Nautilida

Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.

<i>Metacoceras</i> genus of molluscs

Metacoceras is a nautilitoid cephalopod from the Upper Carboniferous (Pennsylvanian) and Permian, the shell of which is moderately evolute with a subquadrate whorl section, bearing nodes on the ventral or umbilical shoulders or both, but otherwise smooth. The siphuncle is small, subcentral and orthochoanitic. The suture has shallow ventral and lateral lobes but no dorsal or annular lobe.

Neoaganides is a small, 1–2 cm diameter subdiscoidal to subglobular goniatitid belonging to the family Pseudohaloritidae that lived from the Late Pennsylvanian to the Late Permian, existing for some 56 million years.

Sosioceras is an Upper Permian shouchangoceratin pseudohaloritid characterized by a subdiscoidal shell, about 2 cm in diameter, marked by growth lines, as with Neoaganides, and having a mature peristome with a double constriction near the periphery. The suture also closely resembles that of Neoaganides.

The Uddenitinae a subfamily of the Medlicottiidae, a family of ammonoid cephalopods included in the Prolecanitida. The Uddenitinae, proposed by Miller and Furnish, and known from the Pennsylvanian and Lower Permian, are transitional between the ancestral Pronoritidae and the more traditional medlicottiids

References

  1. 1 2 The Paleobiology Database 11/17/09
  2. 1 2 3 4 Miller, A.K. and W.M Furnish, 1957, Permian Ammonoids from Southern Arabia; Jour Paleontology V.31, N. 6, pp 1043-1051; Nov. 1957
  3. Goniat on line 11/17/09
  4. Saunders et al. 1999