Pseudohaloritidae Temporal range: | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Cephalopoda |
Subclass: | † Ammonoidea |
Order: | † Goniatitida |
Superfamily: | † Pseudohaloritoidea |
Family: | † Pseudohaloritidae Miller & Furnish, 1957 |
Subfamilies | |
Pseudohaloritidae is the larger of two families that form the goniatitid superfamily Pseudohaloritoidea, the other being the monogenerc Maximitidae. They are part of the vast array of shelled cephalopods known as ammonoids that are more closely related to squids, belemnites, octopuses, and cuttlefish, than to the superficially similar Nautilus . [1]
The Pseudohaloritide which now contains some 14 genera in three subfamilies is characterized by small, subdiscoidal to subglobular, involute shells, the surface of which may be smooth or with coarse longitudinal lirae and/or transverse ribs. The siphuncle is retrosiphonitic, a hold-over character from the nautiloids, usually subcentral or situated within dorsal septal flexure but ventral-marginal in first and second whorls. Sutures have four pairs of lobes, the ventral one being rounded and spatulate, the interior three clustered and narrow. External lobes are either smooth are variably serrate. [1] [2]
The Pseudohaloritidae was established by Miller and Furnish (1957) [2] for three related genera with similar sutures and aberrant siphuncles that are removed from the ventral margin. One, Maximites, has been put in a family of its own, the Maximitidae. The other two, Neoaganides and Pseudohalorites are retained in the Pseudohaloritidae, to which has been added about a dozen more genera, mostly from China. [3]
Maximites differs in having a bifid ventral lobe and a siphuncle that is ventral, but not marginal. It is also older, as early as the Middle Pennsylvanian (m U Carb) rather than from the Upper Pennsylvanian (uU Carb) when pseudohaloritids, as emended, make their first appearance. [4] Maximites is thought to have given rise to the Neoaganides and thus to the Pseudohaloritidae, and in turn may have been derived from Imitoceras. On the other hand, the suture indicates that Neoaganides may have developed directly from Imitoceras. The ventral lobe of Neoaganides is like that of Imitoceras, smooth and rounded, but the lateral lobes differ from those of both Imitoceras and Maximites in being pointed. [2]
The aberrant position of the siphuncle well away from the ventral margin and in some near the dorsum is characteristic of, although not unique to, this family. However no link is indicated between these Pennsylvanian and Permian forms and the Upper Devonian Clymeniids with their well established dorso-marginal siphuncles. [2]
Ammonoids are extinct spiral shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living coleoids than they are to shelled nautiloids. The earliest ammonoids appeared during the Devonian, with the last species vanishing during or soon after the Cretaceous–Paleogene extinction event. They are often called ammonites, which is most frequently used for members of the order Ammonitida, the only remaining group of ammonoids from the Jurassic up until their extinction.
Goniatids, informally goniatites, are ammonoid cephalopods that form the order Goniatitida, derived from the more primitive Agoniatitida during the Middle Devonian some 390 million years ago. Goniatites (goniatitids) survived the Late Devonian extinction to flourish during the Carboniferous and Permian only to become extinct at the end of the Permian some 139 million years later.
Yinoceras is a genus of middle Permian goniatitid ammonite, the type genus for the subfamily Yinoceratinae of the family Pseudohaloritidae.
Dimeroceratidae is one of three families in the Dimeroceratoidea, a goniatid superfamily included in the Ammonoidea; extinct shelled cephalopods with adorally convex septa and usually narrow ventro-marginal siphuncles.
Pseudohaloritoidea, formerly Pseudohaloritaceae, is one of four superfamilies of the goniatitid suborderTornoceratatina. Although attributed the Ruzhencev, 1957 T.J Frest et al. included the Maximitidae and Pseudohaloritidae in the Cheilocerataceae in their May 1981 paper.
Gastrioceratoidea is one of 17 superfamilies in the suborder Goniatitina, ammonoid cephalopods from the Late Paleozoic.
Adrianitidae is a family in the Adrianitaceae, a superfamily of ammonites in the cephalopod order, Goniatitida, known from the Middle Pennsylvanian to the Middle Permian.
Medlicottiidae is a family of ammonoid cephalopods belonging to the Prolecanitida, known from the Upper Carboniferous (Pennsylvanian) to the Early Triassic.
Prolecanitida is an order of extinct ammonoid cephalopods, the major Late Paleozoic group of ammonoids alongside the order Goniatitida. Prolecanitids had narrow shells, discoidal (disc-shaped) to thinly lenticular (lens-shaped). They retained a retrochoanitic siphuncle, a simple form with septal necks extending backwards. As is typical for ammonoids, the siphuncle sits along the ventral margin of the shell.
Prolecanitoidea is a taxonomic superfamily of ammonoids in the order Prolecanitida. Prolecanitoidea is one of two superfamilies in the order, along with the younger and more complex Medlicottioidea. The Prolecanitoidea were a low-diversity and morphologically conservative group. They lived from the Lower Carboniferous up to the Middle Permian. Their shells are generally smooth and discoidal, with a rounded lower edge, a moderate to large umbilicus, and goniatitic to ceratitic sutures. Suture complexity varies from 10 up to 22 total lobes ; new lobes are added from subdivision of saddles adjacent to the original main umbilical lobe.
The Daraelitidae form a family in the ammonoid order Prolecanitida from the Upper Mississippian - Middle Permian characterized by discoidal shells with no prominent sculpture, moderately large umbilicus, and goniatitic or ceratitic sutures with a trifid ventral lobe and few auxiliary lobes.
Permoceras, the sole member of the family Permoceratidae, is a genus of coiled nautiloids with a smooth, compressed involute shell, whorls higher than wide, earlier whorls hidden from view. The venter is rounded as are the ventral and umbilical shoulders, the flanks flattened. The siphuncle is ventrally subcentral. The suture, which is most characteristic, has a deep, narrow pointed ventral lobe and large, asymmetrical pointed lobes on either side.
The Clydonautiloidea are a superfamily within the nautiloid order Nautilida characterized by smooth, generally globular, shells with nearly straight sutures, in early forms, but developing highly differentiated sutures in some later forms. Where known, the siphuncle tends to be central to subcentral.
The Trigonoceratoidea are a superfamily within the Nautilida that ranged from the Devonian to the Triassic, thought to have contained the source for the Nautilaceae in which Nautilus is found.
Grypoceratidae is the longest-lived family of the Trigonoceratoidea, or of the near equivalent Centroceratina; members of the Nautilida from the Upper Paleozoic and Triassic.
The Centroceratidae is the ancestral family of the Trigonoceratoidea and of the equivalent Centroceratina; extinct shelled cephalopods belonging to the order Nautilida
Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.
Neoaganides is a small, 1–2 cm diameter subdiscoidal to subglobular goniatitid belonging to the family Pseudohaloritidae that lived from the Late Pennsylvanian to the Late Permian, existing for some 56 million years.
Shouchangoceras is a genus of goniatitid pseudohaloritid ammonoids and is the type genus for the pseudohaloritid subfamily Shouchangoceratinae. The genus is characterized by a compressed shell as much as 5 cm in diameter, with a strongly constricted mature peristome that has shallow dorsolateral sinus, a moderately deep rounded ventral sinus, but without conspicuous lappets, and ornamented by moderately strong transverse ribs and numerous stronger longitudinal lirae, producing a weakly reticulate pattern. Lobes are attenuate. The siphuncle is within the dorsal septal flecture.
The Uddenitinae a subfamily of the Medlicottiidae, a family of ammonoid cephalopods included in the Prolecanitida. The Uddenitinae, proposed by Miller and Furnish, and known from the Pennsylvanian and Lower Permian, are transitional between the ancestral Pronoritidae and the more traditional medlicottiids.