Rafinesquina Temporal range: | |
---|---|
Rafinesquina ponderosa from the Excello South Outcrop near the Ohio-Kentucky-Indiana tristate area | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Brachiopoda |
Class: | † Strophomenata |
Order: | † Strophomenida |
Family: | † Rafinesquinidae |
Subfamily: | † Rafinesquininae |
Genus: | † Rafinesquina Clarke and Hall, 1892 |
Type species | |
Leptaena alternata Conrad, 1838 | |
Species | |
See Species |
Rafinesquina is an extinct genus of large brachiopod that existed from the Darriwilian to the Ludlow epoch. [1]
The genus was named in honor of polymath Constantine Samuel Rafinesque-Schmaltz. [2]
Rafinesquina's members were epifaunal, meaning they lived on top of the seafloor, not buried within it, and were suspension feeders. [3] Rafinesquina normally have a concavo-convex profile, with radiating striae of alternating size which are crossed with finer concentric striae. [3] Their width is usually greater than their length, like most Strophomenids. Members of this genus had shells that grew in increments, with each increment forming a layer of the shell (much like trees do with their rings). In 1982, Gary D. Rosenberg analyzed specimens of Rafinesquina alternata previously inferred to have lived in a shallow subtidal environment and proposed it could be possible to estimate the total number of days in a lunar month (the period between full moons) during the Late Ordovician using layer counting. [4]
Rafinesquina specimens had a cosmopolitan distribution, and their fossils can be found in the Americas, Europe, and Asia. [1] [5] [6] [7]
Species in the genus Rafinesquina include: [6] [7] [8]
The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Ma) to the start of the Silurian Period 443.8 Mya.
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as being basal lophotrochozoans, or even molluscs.
The Craniidae are a family of brachiopods, the only surviving members of the subphylum Craniiformea. They are the only members of the order Craniida, the monotypic suborder Craniidina, and the superfamily Cranioidea; consequently, the latter two taxa are at present redundant and rarely used.There are three living genera within Craniidae: Neoancistrocrania, Novocrania, and Valdiviathyris. As adults, craniids either live freely on the ocean floor or, more commonly, cement themselves onto a hard object with all or part of the ventral valve.
Strophomenida is an extinct order of articulate brachiopods which lived from the lower Ordovician period to the mid Carboniferous period. Strophomenida is part of the extinct class Strophomenata, and was the largest known order of brachiopods, encompassing over 400 genera. Some of the largest and heaviest known brachiopod species belong to this class. Strophomenids were among the most diverse and abundant brachiopods during the Ordovician, but their diversity was strongly impacted at the Late Ordovician mass extinction. Survivors rediversified into new morphologies in the Silurian, only to be impacted once again at the Late Devonian mass extinction. However, they still survived till the mid Carboniferous.
Brachiopods, phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a stalk-like pedicle projects from an opening near the hinge of one of the valves, known as the pedicle or ventral valve. The pedicle, when present, keeps the animal anchored to the seabed but clear of sediment which would obstruct the opening.
The origin of the brachiopods is uncertain; they either arose from reduction of a multi-plated tubular organism, or from the folding of a slug-like organism with a protective shell on either end. Since their Cambrian origin, the phylum rose to a Palaeozoic dominance, but dwindled during the Mesozoic.
Lingulella is a genus of phosphatic-shelled brachiopod. It is known from the Middle Cambrian Burgess Shale (Canada) to the Upper Ordovician Bromide Formation in North America. 346 specimens of Lingulella are known from the Greater Phyllopod bed, where they comprise 0.66% of the community.
The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.
Concavodonta is an extinct genus of early bivalve in the extinct family Praenuculidae. The genus is one of three genera in the subfamily Concavodontinae. Concavodonta is known solely from late Ordovician, Caradoc epoch, fossils found in Europe and South America. The genus currently contains three accepted species, Concavodonta imbricata, Concavodonta ovalis and the type species Concavodonta ponderata.
Concavodontinae is an extinct subfamily of prehistoric bivalves in the family Praenuculidae. Concavodontinae species lived from the middle Ordovician, Caradoc epoch through the late Ordovician Ashgill epoch. Concavodontinae fossils are found in Europe and South America, and species are thought to have been stationary attached to substrate in shallow infaunal marine water environments where they formed shells of an aragonite composition. The subfamily Concavodontinae was named by Teresa M. Sánchez in 1999.
Paterinata is an extinct class of linguliform brachiopods which lived from the lower Cambrian ("Tommotian") to the Upper Ordovician (Hirnantian). It contains the single order Paterinida and the subfamily Paterinoidea. Despite being some of the earliest brachiopods to appear in the fossil record, paterinides stayed as a relatively subdued and low-diversity group even as other brachiopods diversified later in the Cambrian and Ordovician. Paterinides are notable for their high degree of convergent evolution with rhynchonelliform (articulate) brachiopods, which have a similar set of muscles and hinge-adjacent structures.
Craniopsidae is an extinct family of craniiform brachiopods which lived from the mid-Cambrian to the Lower Carboniferous (Tournaisian). It is the only family in the monotypic superfamily Craniopsoidea and the monotypic order Craniopsida. If one includes the ambiguous Cambrian genus Discinopsis, craniopsids were the first craniiforms to appear, and may be ancestral to craniids and trimerellides. An even earlier Cambrian genus, Heliomedusa, has sometimes been identified as a craniopsid. More recently, Heliomedusa has been considered a stem-group brachiopod related to Mickwitzia.
Strophomenata is an extinct class of brachiopods in the subphylum Rhynchonelliformea.
Siphonotretida is an extinct order of linguliform brachiopods in the class Lingulata. The order is equivalent to the sole superfamily Siphonotretoidea, itself containing the sole family Siphonotretidae. Siphonotretoids were originally named as a superfamily of Acrotretida, before being raised to their own order.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2019.
Askerina is an extinct genus of atrypid brachiopods from subfamily Atrypinae that lived in end-Ordovician (Hirnantian) and Silurian (Aeronian). The type and only known species is Askerina cymbula. Its fossils are known only from the lower parts of the Solvik Formation in central Oslo region, Norway.
Leptaena is an extinct genus of mid-sized brachiopod that existed from the Dariwilian epoch to the Emsian epoch, though some specimens have been found in strata as late in age as the Tournasian epoch. Like some other Strophomenids, Lepteana were epifaunal, meaning they lived on top of the seafloor, not buried within it, and were suspension feeders.
Pugnoides is an extinct genus of brachiopod belonging to the order Rhynchonellida and family Petasmariidae. Specimens have been found in Devonian to Permian beds in North America, Asia, Europe, western Australia, New Zealand,and New Zealand. The genus was particularly widespread in the Visean.
Gertruda Józefa Biernat was a Polish geologist and paleontologist. During World War II, she joined the Polish resistance, participating in the Warsaw Uprising. She is noted for her work in identifying and studying brachiopods of the Ordovician period of the Paleozoic Era, and several have been named in her honor. Biernat worked at institutions of the Polish Academy of Sciences, published major works on brachiopods, participating in paleontological expeditions internationally, including leading a series of expeditions in the Arctic.