Rybushka Formation

Rybushka Formation
Stratigraphic range: Campanian ~84–71  Ma PreꞒ Ꞓ O S D C P T J K Pg N
Type	Geological formation
Lithology
Primary	Sandstone
Other	Phosphorite
Location
Coordinates	51°18′N45°00′E / 51.3°N 45.0°E / 51.3; 45.0
Approximate paleocoordinates	42°36′N40°12′E / 42.6°N 40.2°E / 42.6; 40.2
Region	Penza, Saratov
Country	Russia
Rybushka Formation (Russia)

The Rybushka Formation is a Campanian geologic formation in the Penza and Saratov Oblasts of European Russia. Pterosaur, fish and invertebrate fossils have been recovered from the formation. ^{[1] [2]}

Fossil content

The following fossils have been reported from the formation:

• Amylodon karamysh
• Archaeolamna kopingensis
• Bogolubovia orientalis
• Bonnerichthys gladius ^[3]
• Cretolamna appendiculata
• Ischyodus bifurcatus
• Volgadraco bogolubovi
• Pseudocorax laevis
• Squalicorax kaupi
• Squatina hasei
• Chalmys sp.
• Edaphodon sp.
• Elasmodus sp.
• Eostriatolamia sp.
• Gryphaeostrea sp.
• Heterodontus sp.
• Monticulina sp.
• Prognathodon sp.
• Solariella sp.
• Squatirhina sp.
• ?Clidastes sp.
• Latoplatecarpus willistoni ^[4]
• Chelospharginae indet.
• Elasmobranchii indet.
• Elasmosauridae indet.
• Enchodontidae indet.
• Mosasauridae indet.
• Plesiosauria indet.
• Polycotylidae indet.
• Testudinata indet.
• ?Thoracosaurinae indet.

See also

• List of pterosaur-bearing stratigraphic units

References

1. Sullivan, Robert M.; Lucas, Spencer G.; Spielmann, Justin A. (2011-01-01). Fossil Record 3: Bulletin 53. New Mexico Museum of Natural History and Science.
2. Averianov, Alexander (2014-08-11). "Review of taxonomy, geographic distribution, and paleoenvironments of Azhdarchidae (Pterosauria)". ZooKeys (432): 1–107. Bibcode:2014ZooK..432....1A. doi: 10.3897/zookeys.432.7913 . ISSN   1313-2989. PMC   4141157 . PMID   25152671.
3. Kanarkina, A.; Zverkov, Nikolay G.; Popov, Evgeny V. (2025). "The first record of the pachycormid fish Bonnerichthys in Eurasia marks a global distribution of Late Cretaceous suspension-feeding giants". Journal of Palaeogeography. 100255. doi: 10.1016/j.jop.2025.100255 .
4. Grigoriev, D. V.; Arkhangelsky, M. S.; Zverkov, N. G.; Gubarev, D. I. (2025). "First record of a rare plioplatecarpine mosasaur Latoplatecarpus in Europe as further evidence of a semi-global Campanian marine vertebrate fauna". Cretaceous Research. 106102. doi:10.1016/j.cretres.2025.106102.

Further reading

• M. S. Arkhangelsky, A. O. Averianov, and E. M. Pervushov. 2007. Short-necked plesiosaurs of the Family Polycotylidae from the Campanian of the Saratov Region. Paleontological Journal 41(6):656-660
• A. V. Averianov and E. V. Popov. 2014. A pterosaurian vertebra from the Upper Cretaceous of the Saratov Region. Paleontological Journal 48(3):326-329
• A. O. Averianov, M. S. Arkhangelsky, and E. M. Pervushov. 2008. A new Late Cretaceous azhdarchid (Pterosauria, Azhdarchidae) from the Volga region. Paleontological Journal 42(6):634-642
• A. O. Averianov. 2007. New records of azhdarchids (Pterosauria, Azhdarchidae) from the Late Cretaceous of Russia, Kazakhstan, and Central Asia. Paleontological Journal 41(2):189-197
• N. N. Bogoliubow. 1914. O ozvonkye pterodaktilya iz verkhnemelovykh otlozheniy Saratovskoy gubernii [A pterodactyl vertebra from the Upper Cretaceous deposits of Saratov province]. Ezhegodnikie po Geologii i Mineralogii Rossii/Annuaire Géologique et Minéralogique de la Russie 16(1):1-7