Scolopendromorpha | |
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Scolopendra ovitora guarding her eggs | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Myriapoda |
Class: | Chilopoda |
Subclass: | Pleurostigmomorpha |
Order: | Scolopendromorpha |
Families | |
Scolopendromorpha is an order of centipedes also known as tropical centipedes [1] or bark centipedes. [2] [3] This order includes about 700 species in five families. These centipedes are found nearly worldwide, with tropical and subtropical regions providing the richest diversity in species. [4] This order includes the only known amphibious centipedes, Scolopendra cataracta , Scolopendra paradoxa , and Scolopendra alcyona . [5] [6] [7] [8]
The centipedes in this order are epimorphic, hatching with a full complement of segments. [9] They usually possess 21 or 23 trunk segments with the same number of paired legs. The number of leg pairs is fixed at 21 for most species in this order and fixed at 23 for the remaining species, except for two species with intraspecific variation: Scolopendropsis bahiensis , which has 21 or 23 leg pairs, and S. duplicata , which has 39 or 43 leg pairs. [10] Species in this order have flattened bodies ranging from 9 mm to 30 cm in length. Colors vary but can be vivid sometimes. Antennae are usually gradually attenuated and have from 14 to 34 segments (but usually have 17 to 21 segments). [4]
The order comprises the five families Cryptopidae, Scolopendridae, Mimopidae, Scolopocryptopidae, and Plutoniumidae. Nearly all species in the family Scolopendridae have four ocelli (simple eyes) on each side of the head, and the genus Mimops (family Mimopidae) features a pale area often considered an ocellus on each side of the head, whereas the other three families are blind. [11] [12] Species in the family Scolopocryptopidae have 23 leg-bearing segments, whereas species in all other families in this order have only 21 leg-bearing segments (with the exception of the genus Scolopendropsis in Scolopendridae). [13] [12] [11] [14]
Centipedes are predatory arthropods belonging to the class Chilopoda of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (metameric) creatures with one pair of legs per body segment. All centipedes are venomous and can inflict painful stings, injecting their venom through pincer-like appendages known as forcipules or toxicognaths, which are actually modified legs instead of fangs. Despite the name, no centipede has exactly 100 legs; the number of pairs of legs is an odd number that ranges from 15 pairs to 191 pairs.
Myriapods are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial.
Scolopendridae is a family of large centipedes.
Scolopendra is a species-rich genus of large tropical centipedes of the family Scolopendridae.
Geophilomorpha is an order of centipedes commonly known as soil centipedes. The name "Geophilomorpha" is from Ancient Greek roots meaning "formed to love the earth." This group is the most diverse centipede order, with 230 genera. These centipedes are found nearly worldwide but are absent in Antarctica and most Arctic regions.
The Craterostigmomorpha are the least diverse centipede clade, comprising only two extant species, both in the genus Craterostigmus. Their geographic range is restricted to Tasmania and New Zealand. There is a single ocellus on each side of the head capsule. They have a distinct body plan; their anamorphosis comprises a single stage: in their first moult, they grow from having 12 trunk segments to having 15. Adult centipedes in this order, like those in Scutigeromorpha and Lithobiomorpha, have 15 leg-bearing segments. Their low diversity and intermediate position between the primitive anamorphic centipedes and the derived Epimorpha has led to them being likened to the platypus. They represent the survivors of a once diverse clade. Maternal brooding unites the Craterostigmomorpha with the Epimorpha into the clade Phylactometria which includes Craterostigmomorpha, Scolopendromorpha, and Geophilomorpha. This trait is thought to be closely linked with the presence of sternal pores, which secrete sticky or noxious secretions, which mainly serve to repel predators and parasites. The presence of these pores on the Devonian Devonobius which is included in own order Devonobiomorpha permits its inclusion in this clade, allowing its divergence of Lithobiomorpha from Phylactometria to be dated to 375 million years ago.
Scolopendra morsitans, also known as the Tanzanian blue ringleg or red-headed centipede, is a species of centipede in the family Scolopendridae. S. morsitans is the type species for the genus Scolopendra.
Scolopocryptopidae is a family of blind centipedes in the order Scolopendromorpha. This family includes more than 90 species. Centipedes in this family are found mainly in the Americas and East Asia but have also been recorded in West Africa, New Guinea, and Fiji.
Himantariidae is a monophyletic family of centipedes in the order Geophilomorpha and superfamily Himantarioidea, found almost exclusively in the Northern Hemisphere. Centipedes in this family feature a short head with a concave labral margin bearing a row of denticles, a single dentate lamella and some pectinate lamellae on each mandible, second maxillae with strongly tapering telopodites and slightly spatulate claws, and a stout forcipular segment with short forcipules and a wide tergite; the ultimate legs usually have no pretarsus, and the female gonopods are distinct and biarticulate.
The Cryptopidae are a family of scolopendromorph centipedes. Cryptopids are blind and possess 21 pairs of legs. The genus Cryptops is the numerically largest in the family, comprising over 150 species worldwide.
Dendrothereua is a genus of house centipedes in the family Scutigeridae. There are at least three described species in Dendrothereua, found in the southern United States and the Neotropics.
Scolopendra hardwickei, the Indian tiger centipede, is a species of centipede in the family Scolopendridae.
Mecistocephalidae is a monophyletic family of centipedes in the order Geophilomorpha. It is the only family in the suborder Placodesmata. Most species in this family live in tropical or subtropical regions, but some occur in temperate regions. This family is the third most diverse in the order Geophiliomorpha, with about 170 species, including about 130 species in the genus Mecistocephalus.
Schendylidae is a family of centipedes in the order Geophilomorpha found in the Americas, the Palearctic region, Africa, Madagascar, Australia, and southeast Asia, and also on some Pacific islands.
Scolopocryptops is a genus of bark centipedes in the family Scolopocryptopidae. There are over 20 described species in Scolopocryptops.
Gonibregmatidae are a paraphyletic family of soil centipedes belonging to the superfamily Geophiloidea.
Arthrorhabdus, from the Greek ἄρθρον, a joint, and ῥάβδος, a staff, is a genus of Scolopendrid centipede in the subfamily Scolopendrinae. Species are found in Mexico and the Southern United States, Australia (A. paucispinus & A. mjöbergi), and South Africa (A. formosus). Since a reapprasial in the genus in 2010, the genus only has four species. It may be polyphyletic.
Mecistocephalus is the largest genus of centipedes in the family Mecistocephalidae, with about 140 species. This genus is among the most diverse and widespread of all the genera in the order Geophilomorpha. The British entomologist George Newport first proposed this genus in 1843 to contain a group of centipedes marked by an unusual elongation of the head.
Gonibregmatus is a genus of centipedes in the family Gonibregmatidae. It was described by British entomologist George Newport in 1843.
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