Simocyon

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Simocyon
Temporal range: late Miocene to early Pliocene
Simocyon primigenius.JPG
Simocyon primigenius lower jaw at Musee d'Histoire Naturelle Paris.
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Ailuridae
Subfamily: Simocyoninae
Genus: Simocyon
Wagner, 1858
Species
  • S. batalleri(Viret, 1929)
  • S. diaphorus(Kaup, 1832)
  • ?S. hungaricusKretzoi in Kadic and Kretzoi, 1927
  • S. primigenius(type species) (Roth and Wagner, 1854)
Synonyms
  • AmphalopexKaup, 1861
  • AraeocyonThorpe, 1922
  • Metarctos Gaudry, 1860

Simocyon ("short-snouted dog") is a genus of extinct carnivoran mammal in the family Ailuridae. Simocyon, which was about the size of a mountain lion, lived in the late Miocene and early Pliocene epochs, and has been found in Europe, Asia, and rarely, North America [1] and Africa. [2]

Contents

Classification

Reconstructed skull and head Simocyon batalleri head.jpg
Reconstructed skull and head

The relationship of Simocyon to other carnivores has been controversial, but studies of the structure of its ear, teeth, and ankle now indicate that its closest living relative is the red panda, Ailurus , [3] [1] although it is different enough to be classified in a separate subfamily (Simocyoninae) along with related genera Alopecocyon and Actiocyon. While the red panda is primarily herbivorous, the teeth and skull of Simocyon indicate that it was carnivorous, and it may have engaged in some bone-crushing, like living hyenas. [1] . The skeleton of Simocyon indicates that, like the red panda, it could climb trees, although it probably also spent considerable time on the ground. [4] Simocyon and Ailurus both have a radial sesamoid, an unusual bone in the wrist that acts as a false thumb. [5] Its competitors during its time period were ailuropodine and tremarctine bears, nimravid false cats, and early canids and felids.

Related Research Articles

<span class="mw-page-title-main">Red panda</span> Species of mammal in Asia

The red panda, also known as the lesser panda, is a small mammal native to the eastern Himalayas and southwestern China. It has dense reddish-brown fur with a black belly and legs, white-lined ears, a mostly white muzzle and a ringed tail. Its head-to-body length is 51–63.5 cm (20.1–25.0 in) with a 28–48.5 cm (11.0–19.1 in) tail, and it weighs between 3.2 and 15 kg. It is well adapted to climbing due to its flexible joints and curved semi-retractile claws.

<span class="mw-page-title-main">Sesamoid bone</span> Bone embedded within a tendon or muscle

In anatomy, a sesamoid bone is a bone embedded within a tendon or a muscle. Its name is derived from the Greek word for 'sesame seed', indicating the small size of most sesamoids. Often, these bones form in response to strain, or can be present as a normal variant. The patella is the largest sesamoid bone in the body. Sesamoids act like pulleys, providing a smooth surface for tendons to slide over, increasing the tendon's ability to transmit muscular forces.

<span class="mw-page-title-main">Amphicyonidae</span> Extinct family of carnivores

Amphicyonidae is an extinct family of terrestrial carnivorans belonging to the suborder Caniformia. They first appeared in North America in the middle Eocene, spread to Europe by the late Eocene, and further spread to Asia and Africa by the early Miocene. They had largely disappeared worldwide by the late Miocene, with the latest recorded species at the end of the Miocene in Africa. They were among the first carnivorans to evolve large body size. Amphicyonids are colloquially referred to as "bear-dogs".

<span class="mw-page-title-main">Nimravidae</span> Extinct family of mammals in the order Carnivora

Nimravidae is an extinct family of carnivorans, sometimes known as false saber-toothed cats, whose fossils are found in North America and Eurasia. Not considered to belong to the true cats, the nimravids are generally considered closely related and classified as a distinct family in the suborder Feliformia. Fossils have been dated from the Middle Eocene through the Late Miocene epochs, spanning about 33.2 million years.

<span class="mw-page-title-main">Ailuridae</span> Family of carnivores

Ailuridae is a family in the mammal order Carnivora. The family consists of the red panda and its extinct relatives.

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Megistotherium is an extinct genus of hyaenodont belonging to the family Hyainailouridae that lived in Africa.

<i>Amphicyon</i> Extinct genus of carnivores

Amphicyon is an extinct genus of large carnivorans belonging to the family Amphicyonidae, subfamily Amphicyoninae, from the Miocene epoch. Members of this family received their vernacular name for possessing bear-like and dog-like features. They ranged over North America, Europe, Asia, and Africa.

<span class="mw-page-title-main">Barbourofelidae</span> Extinct family of feliform carnivorans

Barbourofelidae is an extinct family of carnivorans of the suborder Feliformia, sometimes known as false saber-toothed cats, that lived in North America, Eurasia and Africa during the Miocene epoch and existed for about 7.9 million years. Thought to be an independent lineage from the Nimravidae and Machairodontinae, which had all attained elongated canines, recent research argues that it may be a subfamily of the Nimravidae, extending its biochronological range into the Miocene, although this issue is not yet fully resolved.

<span class="mw-page-title-main">Ursoidea</span> Superfamily of mammals

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Alopecocyon is a fossil genus of ailurid belonging to the subfamily Simocyoninae. It has two species, Alopecocyon goeriachensis and Alopecocyon getti. It is based on fragmentary fossils dating to the middle Miocene of both Europe and Asia.

<i>Palaearctonyx</i> Extinct genus of carnivores

Palaearctonyx is an extinct genus of omnivorous placental mammals from clade Carnivoraformes, that lived in North America from early to middle Eocene.

<i>Agnotherium</i> Extinct genus of carnivores

Agnotherium is a genus of large sized carnivoran mammals, belonging to the Amphicyonidae, which has been found in Western Europe, and possibly China and Northern Africa, and lived during the Late Miocene epoch. Despite only being known from fragmentary remains, the genus notable for hypercarnivorous adaptions, which have been said to represent the "apex" among its family.

<span class="mw-page-title-main">Cerro de los Batallones</span> Eye to Ancient Age

Cerro de los Batallones is a hill at Torrejón de Velasco, Madrid, Spain where a number of fossil sites from the Upper Miocene (MN10) have been found. Nine sites have been discovered with predominantly vertebrate fossils, invertebrates and plants being less represented. The first deposits were discovered accidentally in July 1991.

Parailurus is a genus of extinct carnivoran mammal in the family Ailuridae. It was about 50% larger than Ailurus and lived in the early to late Pliocene Epoch, and its fossils have been found in Europe, North America, and Japan.

Sivanasua is an extinct genus of carnivorous mammal found across Miocene Europe including Germany, Austria, France, Spain and the Czech Republic. Like other lophocyonids, Sivanasua had unusual lophodont dentition, meaning the molars had ridges across the grinding surface of the molars, an adaptation believed to be indicative of a herbivorous diet.

<span class="mw-page-title-main">Simocyoninae</span> Extinct subfamily of carnivores

Simocyoninae is an extinct subfamily of Ailuridae. The taxonomic history of this group was complicated, as researchers placed various fossil caniform genera into the subfamily. In addition to Simocyon, there was also Oligobunis, Cephalogale, and Enhydrocyon. This subfamily was initially classified within the Canidae or dog family. This idea went even further in 1910 when American paleontologist Henry Fairfield Osborn considered dholes, African wild dogs, and bush dogs to be the only extant representation left of the simocyonines. This was in large part to the overall similarity in the morphology of their molars, which suggested a shared ancestry of hypercarnivory. This view point was not supported by European paleontologists who believed that Simocyon was more closely related to musteloids. Soon the Simocyoninae were found as extinct subfamily of procyonids, or members of the raccoon-family, due to similarity of the basicranium of these animals. This was supported by subsequent researchers. It was not until a more completed skull of a Simocyon found in north Shaanxi, China was described by Wang (1997) who found that Simocyoninae is closely related Ailurinae on the basis of cranial and dental characteristics.

<span class="mw-page-title-main">Thaumastocyoninae</span> Extinct subfamily of amphicyonids

Thaumastocyoninae is an extinct subfamily of amphicyonids, large terrestrial carnivores, which inhabited what is now Europe during the Miocene epoch. The subfamily was erected by Hürzeler (1940), and is defined by the complete suppression of m1 metaconid, reduction of the premolars, except the p4, which is reinforced, and the oblique abrasion of the teeth, and the possession of hypercarnivorous tendencies. Thaumastocyonines are poorly known, with only about 65 dental specimens, most of those isolated teeth, being known as of 2020, although more complete remains have recently been discovered.

<i>Actiocyon</i> Extinct genus of carnivores

Actiocyon was an extinct genus of ailurid that existed from the Middle Miocene of North America. The type species, Actiocyon leardi, was described in 1947 by the American paleontologist Chester Stock.

Namibiocyon is an extinct genus of carnivoran mammals, belonging to the family Amphicyonidae, that lived in Namibia during the Early Miocene epoch. Before the erection of this taxon in 2022, the type and only species, N. ginsburgi, had been assigned to a variety of other genera. It is notable for its adaptions toward hypercarnivory.

Protursus is a fossil genus of ailurid with a single species Protursus simpsoni, known from a single left m2 tooth.

References

  1. 1 2 3 Peigné, S.; Salesa, M. J.; Antón, M.; Morales, J. (2005). "Ailurid carnivoran mammal Simocyon from the late Miocene of Spain and the systematics of the genus" (PDF). Acta Palaeontologica Polonica. 50 (2): 219–238.
  2. Howell, F. Clark; Garcia, Nuria (December 2007). "Carnivora (Mammalia) From Lemudong'o (Late Miocene: Narok District, Kenya)" (PDF). Kirtlandia. Cleveland: Cleveland Museum of Natural History. 556: 121–139. Archived from the original (PDF) on 2016-03-04. Retrieved 2009-10-15.
  3. Wang, Xiaoming (1997). "New cranial material of Simocyon from China, and its implications for phylogenetic relationship to the red panda(Ailurus)". Journal of Vertebrate Paleontology. 17 (1): 184–198. Bibcode:1997JVPal..17..184W. doi:10.1080/02724634.1997.10010963.
  4. Salesa, M. J.; Antón, M.; Peigné, S.; Morales, J. (2008). "Functional anatomy and biomechanics of the postcranial skeleton of Simocyon batalleri (Viret, 1929) (Carnivora, Ailuridae) from the late Miocene of Spain". Zoological Journal of the Linnean Society. 152 (3): 593–621. doi:10.1111/j.1096-3642.2007.00370.x.
  5. Salesa, Manuel J.; Antón, Mauricio; Peigné, Stéphane; Morales, Jorge (2006). "Evidence of a false thumb in a fossil carnivore clarifies the evolution of pandas". Proceedings of the National Academy of Sciences. 103 (2): 379–382. Bibcode:2006PNAS..103..379S. doi: 10.1073/pnas.0504899102 . PMC   1326154 . PMID   16387860.

Bibliography