Snow camouflage

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Ptarmigan, changing colour from winter to disruptively patterned summer camouflage in springtime. The male is still mostly in winter plumage. Rock Ptarmigan (Lagopus Muta).jpg
Ptarmigan, changing colour from winter to disruptively patterned summer camouflage in springtime. The male is still mostly in winter plumage.
Red Army soldiers in snow camouflage near Moscow, December 1941. There are at least 11 soldiers in the image. RIAN archive 284 The war in winter.jpg
Red Army soldiers in snow camouflage near Moscow, December 1941. There are at least 11 soldiers in the image.

Snow camouflage is the use of a coloration or pattern for effective camouflage in winter, often combined with a different summer camouflage. Summer patterns are typically disruptively patterned combinations of shades of browns and greys, up to black, while winter patterns are dominated by white to match snowy landscapes.

Contents

Among animals, variable snow camouflage is a type of seasonal polyphenism with a distinct winter plumage or pelage. It is found in birds such as the rock ptarmigan, lagomorphs such as the Arctic hare, mustelids such as the stoat, and one canid, the Arctic fox. Since these have evolved separately, the similar appearance is due to convergent evolution. This was used as early evidence for natural selection. Some high Arctic species like the snowy owl and polar bear however remain white all year round.

In military usage, soldiers often either exchange their disruptively-patterned summer uniforms for thicker snow camouflage uniforms printed with mainly-white versions of camouflage patterns in winter, or they wear white overalls over their uniforms. Some armies have made use of reversible uniforms, printed in different seasonal patterns on their two sides. Vehicles and guns are often simply repainted in white. Occasionally, aircraft too are repainted in snow camouflage patterns.

Among animals

White as camouflage

The snowy owl remains white all year round. A Snowy Owl in Flight David Hemmings.jpg
The snowy owl remains white all year round.

Charles Darwin mentioned the white winter coloration of the ptarmigan in his 1859 Origin of Species : [1]

When we see ... the alpine ptarmigan white in winter, the red-grouse the colour of heather, and the black-grouse that of peaty earth, we must believe that these tints are of service to these birds ... in preserving them from danger. [1]

The white protective coloration of arctic animals was noted by an early student of camouflage, the naturalist Alfred Russel Wallace, in his 1889 book Darwinism ; he listed the polar bear, the American polar hare, the snowy owl and the gyr falcon as remaining white all year, while the arctic fox, arctic hare, ermine and ptarmigan change their colour, and observed "the obvious explanation", that it was for concealment, at a time when Darwinism was at a low ebb. [2] Later zoologists such as Hugh B. Cott have echoed his observations, adding that other animals of the Arctic such as musk ox, moose, reindeer, wolverine and raven never become white "even in the coldest parts of their range". Cott noted that both animals that hunt, like polar bear and stoat, and prey animals like ptarmigan and mountain hare, require camouflage to hide from prey or from predators respectively. [3] There is little experimental evidence for the adaptiveness of white as camouflage, though the ornithologist W. L. N. Tickell, reviewing proposed explanations of white plumage in birds, writes that in the ptarmigan "it is difficult to escape the conclusion that cryptic brown summer plumage becomes a liability in snow, and white plumage is therefore another cryptic adaptation." All the same, he notes, "in spite of winter plumage, many Ptarmigan in NE Iceland are killed by Gyrfalcons throughout the winter." [4]

Seasonal polyphenism

Willow grouse of Northern forests and tundra change plumage in winter; British populations (red grouse) do not. Denali National Park Ptarmigan.jpg
Willow grouse of Northern forests and tundra change plumage in winter; British populations (red grouse) do not.

Some animals of the far north, like the snowshoe and Arctic hares, Arctic fox, stoat, and rock ptarmigan change their coat colour (by moulting and growing new fur or feathers) from brown or grey summer camouflage to white in the winter; the Arctic fox is the only species in the dog family to do so. [5] However, Arctic hares which live in the far north of Canada, where summer is very short, remain white year-round. [5] [6] Since these animals in widely separated groups have evolved separately, the similarity of coloration is due to convergent evolution, on the presumption that natural selection favours a particular coloration in a particular environment. [7]

The seasonal polyphenism in willow grouse differs between Scottish and Scandinavian populations. In Scotland, grouse have two plumages (breeding and non-breeding), while in Scandinavia there is a third plumage, a white winter morph. The genetic basis for this is not in the melanin pigment system, and is probably due to regulatory changes. [8] The behaviour of moulting females in springtime depends on their plumage state: they tend to sit on snow while they are mainly white, but choose the border between bare ground and snow when they have more dark feathers. They seem to be choosing the best compromise between camouflage and food quality. [9]

The effects of climate change can lead to a mismatch between the seasonal coat coloration of arctic animals such as snowshoe hares with the increasingly snow-free landscape. [10]

Seasonal polyphenism in animals
SpeciesIn summerIn winter
Snowshoe hare Lepus americanus 5459 cropped.jpg Snowshoe Hare, Shirleys Bay.jpg
Arctic fox Arcticfox-3.jpg Iceland-1979445 (cropped).jpg
Stoat Mustela erminea.jpg Mustela erminea winter cropped.jpg
Rock ptarmigan Alpenschneehuhn auf der Totalpe, Lagopus muta 1.JPG Ptarmigan on Beinn Chuirn - geograph.org.uk - 339986.jpg

Military usage

Austro-Hungarian ski patrol on Italian front in snow camouflage, 1915-1918 Austro-Hungarian ski patrol on Italian front in snow camouflage 1915-1918.jpg
Austro-Hungarian ski patrol on Italian front in snow camouflage, 1915-1918

The principle of varying coloration with the changing seasons has military applications.

First World War

In the First World War, firing and observation positions were hand-painted in disruptive patterns by artists known as camoufleurs, and they sometimes varied their patterns seasonally. Uniforms were largely of a single colour, such as the British khaki; [11] but snow camouflage clothes came into use in some armies as the war progressed. [12] [13] For example, Austro-Hungarian troops on the Italian front used skis and wore snow camouflage smocks and overtrousers over their uniforms, and improvised white cloths over their uniform caps. [14] Also French mountain infantry and other, specialized units in Imperial Russian, Turkish and German armies adopted these.

Second World War

Several armies in the Second World War in Northern European countries preferred separate winter uniforms rather than oversuits. The Waffen-SS went a step further, developing reversible uniforms with separate schemes for summer and autumn, as well as white winter oversuits. Other German units fighting in Eastern Europe were at first poorly equipped for winter, having to make do with ordinary summer uniforms, but in the winter of 1942 to 1943 new white two-piece hooded oversuits with long mitten gauntlets started to arrive. American troops in Europe in the winter of 1944 to 1945 improvised snow capes and helmet covers from white cloth such as bed linen. [15] [16] [17]

The Red Army issued a report, "Tactical and Technical trends, No. 17" in January 1943 on the camouflage of tanks in winter. It advised either all-white using zinc white or titanium white paint for level, open country, or disruptive two-colour winter camouflage for areas with more variety including "forests, underbrush, small settlements, thawed patches of earth". The two colours could be achieved either by leaving around a quarter to a third of the vehicle's summer camouflage uncovered, or by repainting the whole vehicle in white with dark gray or gray-brown spots. Units were advised not to paint all their vehicles identically, but to have some tanks all white, some in white with green stripes, and some in white and gray or gray-brown. Winter camouflage was not limited to paint: tracks left in the snow were to be obliterated, vehicles parked in cover, headlights covered with white fabric, shelters constructed, or else vehicles covered with white fabric, or dark fabric scattered with snow. [18]

Seasonal camouflage in the Second World War
ExampleIn summerIn winter
2-pounder British anti-tank gun
Summer crew in British Battledress
winter crew in improvised camouflage
The British Army in the Middle East 1942 E9610.jpg The British Army in France 1940 F2606.jpg
Red Army soldiers
Hooded winter overalls
Saur-Mogila Soviet soldiers colorized.png Bundesarchiv Bild 183-E0406-0022-001, Russland, Kesselschlacht Stalingrad.jpg
Finnish artillery
Improvised winter camouflage
KTR15 tykki.jpg Finnish-artillery-76k02-1902-winterwar.png
Wehrmacht/Waffen-SS soldiers and armour
Two-part winter uniforms
Bundesarchiv Bild 101I-022-2948-23, Russland, Panzer VI (Tiger I), Munition Recolored.jpg Bundesarchiv Bild 101III-Roth-173-01, Russland, Raum Charkow, Jagdpanzer.jpg

Post-war

Later in the 20th century, as disruptively patterned uniforms became widespread, winter variants of patterns such as Flecktarn were developed, the background colour (such as green) being replaced with white to form a Schneetarn (snow camouflage pattern). [19] Aircraft deployed in winter have sometimes been snow camouflaged, as with Sepecat Jaguars on exercise in Norway. [20]

Seasonal camouflage since the Second World War
ExampleIn summerIn winter
Flecktarn/Schneetarn
disruptive camouflage patterns
Flecktarn.jpg M84Snow.jpg
Finnish Defence Forces
digital desert/snow patterns
M04camo.jpg M05 snow pattern.jpg
Royal Air Force Jaguar
summer/winter paint schemes
Jaguar 54 Sqn China Lake CA 1999.jpg Sepecat Jaguar GR1A, UK - Air Force AN1985498.jpg
Ghillie suit
summer/winter
Scharfschutze (25393720634).jpg NATO ProAtpo Ghillie Suit Italian Alpine troops barret 50.cal.jpg

Related Research Articles

<span class="mw-page-title-main">Camouflage</span> Concealment in plain sight by any means, e.g. colour, pattern and shape

Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the battledress of a modern soldier, and the leaf-mimic katydid's wings. A third approach, motion dazzle, confuses the observer with a conspicuous pattern, making the object visible but momentarily harder to locate, as well as making general aiming easier. The majority of camouflage methods aim for crypsis, often through a general resemblance to the background, high contrast disruptive coloration, eliminating shadow, and countershading. In the open ocean, where there is no background, the principal methods of camouflage are transparency, silvering, and countershading, while the ability to produce light is among other things used for counter-illumination on the undersides of cephalopods such as squid. Some animals, such as chameleons and octopuses, are capable of actively changing their skin pattern and colours, whether for camouflage or for signalling. It is possible that some plants use camouflage to evade being eaten by herbivores.

<span class="mw-page-title-main">Arctic hare</span> Species of mammal

The Arctic hare is a species of hare highly adapted to living in the Arctic tundra and other icy biomes. The Arctic hare survives with shortened ears and limbs, a small nose, fat that makes up close to 20% of its body, and a thick coat of fur. It usually digs holes in the ground or under the snow to keep warm and to sleep. Arctic hares look like rabbits but have shorter ears, are taller when standing, and, unlike rabbits, can thrive in extreme cold. They can travel together with many other hares, sometimes huddling with dozens or more, but are usually found alone, sometimes taking more than one partner. The Arctic hare can run up to 60 kilometres per hour (40 mph).

<span class="mw-page-title-main">Willow ptarmigan</span> Species of bird

The willow ptarmigan is a bird in the grouse subfamily Tetraoninae of the pheasant family Phasianidae. It is also known as the willow grouse and in Ireland and Britain, where the subspecies L. l. scotica was previously considered to be a separate species, as the red grouse. It breeds in birch and other forests and moorlands in northern Europe, the tundra of Scandinavia, Siberia, Alaska and Canada, in particular in the provinces of Newfoundland and Labrador and Quebec. It is the state bird of Alaska. In the summer the birds are largely brown, with dappled plumage, but in the winter they are white with some black feathers in their tails. The species has remained little changed from the bird that roamed the tundra during the Pleistocene. Nesting takes place in the spring when clutches of four to ten eggs are laid in a scrape on the ground. The chicks are precocial and soon leave the nest. While they are young, both parents play a part in caring for them. The chicks eat insects and young plant growth while the adults are completely herbivorous, eating leaves, flowers, buds, seeds and berries during the summer and largely subsisting on the buds and twigs of willow and other dwarf shrubs and trees during the winter.

<span class="mw-page-title-main">Rock ptarmigan</span> Species of bird

The rock ptarmigan is a medium-sized game bird in the grouse family. It is known simply as the ptarmigan in Europe. It is the official bird for the Canadian territory of Nunavut, where it is known as the aqiggiq (ᐊᕿᒡᒋᖅ), and the official game bird for the province of Newfoundland and Labrador. In Japan, it is known as the raichō (雷鳥), which means "thunder bird". It is the official bird of Gifu, Nagano, and Toyama Prefectures and is a protected species nationwide. Unlike many arctic bird species, ptarmigan do not gain substantial mass to hibernate over winter.

<span class="mw-page-title-main">Rough-legged buzzard</span> Species of bird

The rough-legged buzzard (Europe) or rough-legged hawk is a medium-large bird of prey. It is found in Arctic and Subarctic regions of North America, Europe, and Russia during the breeding season and migrates south for the winter. It was traditionally also known as the rough-legged falcon in such works as John James Audubon's The Birds of America.

<span class="mw-page-title-main">White-tailed ptarmigan</span> Species of bird

The white-tailed ptarmigan, also known as the snow quail, is the smallest bird in the grouse family. It is a permanent resident of high altitudes on or above the tree line and is native to Alaska and the mountainous parts of Canada and the western United States. Its plumage is cryptic and varies at different times of the year. In the summer it is speckled in gray, brown and white whereas in winter it is wholly white. At all times of year the wings, belly and tail are white. The white-tailed ptarmigan has a diet of buds, leaves, flowers and seeds. The nest is a simple depression in the ground in which up to eight eggs are laid. After hatching, the chicks soon leave the nest. At first they eat insects but later move on to an adult diet, their mother using vocalisations to help them find suitable plant food. The population seems to be stable and the IUCN lists this species as being of "Least Concern".

<i>Lagopus</i> Genus of birds

Lagopus is a small genus of birds in the grouse subfamily commonly known as ptarmigans. The genus contains three living species with numerous described subspecies, all living in tundra or cold upland areas.

<span class="mw-page-title-main">Military camouflage</span> Camouflage used to protect from enemy observation

Military camouflage is the use of camouflage by an armed force to protect personnel and equipment from observation by enemy forces. In practice, this means applying colour and materials to military equipment of all kinds, including vehicles, ships, aircraft, gun positions and battledress, either to conceal it from observation (crypsis), or to make it appear as something else (mimicry). The French slang word camouflage came into common English usage during World War I when the concept of visual deception developed into an essential part of modern military tactics. In that war, long-range artillery and observation from the air combined to expand the field of fire, and camouflage was widely used to decrease the danger of being targeted or to enable surprise. As such, military camouflage is a form of military deception in addition to cultural functions such as political identification.

<span class="mw-page-title-main">Hugh B. Cott</span> English zoologist and camouflage expert (1900–1987)

Hugh Bamford Cott was a British zoologist, an authority on both natural and military camouflage, and a scientific illustrator and photographer. Many of his field studies took place in Africa, where he was especially interested in the Nile crocodile, the evolution of pattern and colour in animals. During the Second World War, Cott worked as a camouflage expert for the British Army and helped to influence War Office policy on camouflage. His book Adaptive Coloration in Animals (1940), popular among serving soldiers, was the major textbook on camouflage in zoology of the twentieth century. After the war, he became a Fellow of Selwyn College, Cambridge. As a Fellow of the Zoological Society of London, he undertook expeditions to Africa and the Amazon to collect specimens, mainly reptiles and amphibians.

<span class="mw-page-title-main">Countershading</span> Camouflage to counteract self-shading

Countershading, or Thayer's law, is a method of camouflage in which an animal's coloration is darker on the top or upper side and lighter on the underside of the body. This pattern is found in many species of mammals, reptiles, birds, fish, and insects, both in predators and in prey.

<span class="mw-page-title-main">Animal coloration</span> General appearance of an animal

Animal colouration is the general appearance of an animal resulting from the reflection or emission of light from its surfaces. Some animals are brightly coloured, while others are hard to see. In some species, such as the peafowl, the male has strong patterns, conspicuous colours and is iridescent, while the female is far less visible.

<span class="mw-page-title-main">Fur</span> Soft, thick, hairy coat of a mammal

Fur is a thick growth of hair that covers the skin of almost all mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket that keeps the animal warm.

<span class="mw-page-title-main">Disruptive coloration</span> Camouflage to break up an objects outlines

Disruptive coloration is a form of camouflage that works by breaking up the outlines of an animal, soldier or military vehicle with a strongly contrasting pattern. It is often combined with other methods of crypsis including background colour matching and countershading; special cases are coincident disruptive coloration and the disruptive eye mask seen in some fishes, amphibians, and reptiles. It appears paradoxical as a way of not being seen, since disruption of outlines depends on high contrast, so the patches of colour are themselves conspicuous.

<i>Adaptive Coloration in Animals</i> 1940 textbook on camouflage, mimicry and aposematism by Hugh Cott

Adaptive Coloration in Animals is a 500-page textbook about camouflage, warning coloration and mimicry by the Cambridge zoologist Hugh Cott, first published during the Second World War in 1940; the book sold widely and made him famous.

<i>Concealing-Coloration in the Animal Kingdom</i> Book by Abbott Handerson Thayer

Concealing-Coloration in the Animal Kingdom: An Exposition of the Laws of Disguise Through Color and Pattern; Being a Summary of Abbott H. Thayer's Discoveries is a book published ostensibly by Gerald H. Thayer in 1909, and revised in 1918, but in fact a collaboration with and completion of his father Abbott Handerson Thayer's major work.

<i>Animal Coloration</i> (book) 1892 book by Frank Evers Beddard

Animal Coloration, or in full Animal Coloration: An Account of the Principal Facts and Theories Relating to the Colours and Markings of Animals, is a book by the English zoologist Frank Evers Beddard, published by Swan Sonnenschein in 1892. It formed part of the ongoing debate amongst zoologists about the relevance of Charles Darwin's theory of natural selection to the observed appearance, structure, and behaviour of animals, and vice versa.

<span class="mw-page-title-main">Coloration evidence for natural selection</span> Early evidence for Darwinism from animal coloration

Animal coloration provided important early evidence for evolution by natural selection, at a time when little direct evidence was available. Three major functions of coloration were discovered in the second half of the 19th century, and subsequently used as evidence of selection: camouflage ; mimicry, both Batesian and Müllerian; and aposematism.

<span class="mw-page-title-main">Distractive markings</span>

Distractive markings serve to camouflage animals or military vehicles by drawing the observer's attention away from the object as a whole, such as noticing its outline. This delays recognition. The markings necessarily have high contrast and are thus in themselves conspicuous. The mechanism therefore relies, as does camouflage as a whole, on deceiving the cognition of the observer, not in blending with the background.

References

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