Black grouse | |
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Male | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Galliformes |
Family: | Phasianidae |
Genus: | Lyrurus |
Species: | L. tetrix |
Binomial name | |
Lyrurus tetrix (Linnaeus, 1758) | |
Black grouse range [2] | |
Distribution in Europe. [2] | |
Synonyms [3] | |
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The black grouse (Lyrurus tetrix), also known as northern black grouse, Eurasian black grouse, blackgame or blackcock, [4] is a large game bird in the grouse family. It is a sedentary species, spanning across the Palearctic in moorland and steppe habitat when breeding, often near wooded areas. They will spend the winter perched in dense forests, feeding almost exclusively on the needles of conifers. The black grouse is one of two species of grouse in the genus Lyrurus , the other being the lesser-known Caucasian grouse.
The female is greyish-brown and has a cackling or warbling call. She takes all responsibility for nesting and caring for the chicks, as typical with most galliforms.
The black grouse's genome was sequenced in 2014. [5]
The black grouse was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae under the binomial name Tetrao tetrix. [6] Both Tetrao and tetrix come from Ancient Greek words referring to some form of game bird. [7] The black grouse is now placed in the genus Lyrurus that was introduced in 1832 by the English naturalist William John Swainson. [8] [9]
The male and female are sometimes referred to by their folk names, blackcock and greyhen, respectively. These names first occur in the literature with John Ray in 1674. [10] Heathcock and Heathhen are also common names. [11]
The black grouse has six recognized subspecies. [12] Black grouse populations differ slightly in size and coloration, with birds increasing in size further east of their range: [13]
The black grouse is a large bird with males measuring roughly around 60 centimetres (24 in) in length and weighing 1,100–1,250 grams (2.43–2.76 lb), sometimes up to 2,100 grams (4.6 lb), with females approximately 45 cm (18 in) and weighing 750–1,100 grams (1.65–2.43 lb). [13] The cock's fancy plumage is predominantly black with deep-blue hues on his neck and back, which contrasts the white wingline and undertail coverts, as well as red bare skin above each eye. On the other hand, the hen is much drabber and cryptically colored to blend in easily with the dense undergrowth, especially when nesting. The black grouse, along with the Caucasian grouse, has long outer rectrices (tail feathers) that curl outward and arranged in a way it resembles the frame of a Greek lyre, hence the genus name, Lyrurus.
Black grouse can be found on open habitats across Europe (Swiss-Italian-French Alps especially) from Great Britain through Scandinavia, Estonia and across Russia. Although believed to once to live in Ireland, it now no longer resides there. In Eastern Europe they can be found in Hungary, Latvia, Lithuania, Poland, Belarus, Romania and Ukraine. There is a population in the Alps, and isolated remnants in Germany, France, Belgium and the Netherlands. [14] It formerly occurred in Denmark, but the Danish Ornithological Society (DOF) has considered it extinct since 2001. The species disappeared from Bulgaria in the 19th century. [15] In Asia, a huge portion of their population can be found in Russia (particularly southern Siberia), though they also inhabit parts of Kazakhstan, Mongolia, China, and possibly Korea. [16]
Black grouse are adapted to an extensive array of habitats across Eurasia, though most frequently utilize the transitionary zones between forests and open clearings, especially steppe, heathland, grassland and pasture when near agricultural fields. Depending on the season, they will overwinter in large flocks in dense forests, and feed primarily on the leaves and buds of coniferous and broadleaf trees, such as Scots pine, Siberian larch, silver birch, and Eurasian aspen. Throughout the spring and summer, they tend to favor open spaces to seek potential mates and raise broods, switching their diet to berries, shoots and stems of cranberries, bog bilberries, myrtleberries, and other Vaccinium shrubs. [13] They avoid the most extreme of desert and polar regions.
This section needs additional citations for verification .(September 2010) |
Although this species has declined throughout most of its range in western Europe, it is not considered to be vulnerable globally due to the large population (global estimate is 15–40 million individuals) and slow rate of decline. [1] Its decline is due to loss of habitat, disturbance, predation by foxes, crows, etc., and small populations gradually dying out.[ citation needed ]
The IUCN implemented a Black Grouse Action Plan 2007–2010. This has looked at local populations that are vulnerable to the extinction vortex. For example, in Styria, Austria.[ citation needed ]
In the United Kingdom black grouse are found in upland areas of Wales, the Pennines and most of Scotland. Best looked for on farmland and moorland with nearby forestry or scattered trees. They have traditional lek sites where the males display.[ citation needed ]
They have declined in some parts of the UK (especially England), having disappeared from many of their former haunts. They are now extirpated in Lancashire, Derbyshire, Exmoor, East Yorkshire, New Forest, Nottinghamshire, Worcestershire, Quantock Hills, Cornwall, Dartmoor, Kent, Wiltshire and Surrey.[ citation needed ]
A programme to re-introduce black grouse into the wild started in 2003 in the Upper Derwent Valley area of the Peak District in England. 30 grouse were released in October 2003, followed by 10 male grouse in December 2004 and a further 10 males and 10 females in April 2005. The programme is being run jointly by the National Trust, Severn Trent Water and Peak District National Park.[ citation needed ]
Conservation groups helping to revive the black grouse include the RSPB and the Game & Wildlife Conservation Trust.[ citation needed ]
In France there has been much work regarding the birds and their risk of flying into ski-lifts. [17]
From 1950 to 2000, local black grouse populations have steadily diminished in Manchuria and northeastern China by about 39%, with birds being most affected (and possibly extirpated) in the Jilin Province. Though the exact causes for local declines remain largely unclear, habitat loss and excess hunting have played a part in the overall reduction. Shifts in their location, however can happen concurrently with the depletion over decades, so short-term research projects may not provide much proof of driving forces behind population alterations. [18]
Based on historical info gathered from various sources (including wildlife surveys and scientific papers), aspen, birch, and poplar make up the black grouse's staple diet and habitat. The climate and precipitation during the month of June is also linked to the breeding success of black grouse. [18]
Black grouse have a very distinctive and well-recorded courtship ritual. Every dawn in the spring, male grouse begin competition with other males in hopes of attracting a hen to mate with. They will display to signal their territory and vigor by fanning out their elaborate lyre-shaped tails and inflating their necks on designated open ground called a lek. Their song consists of a long, dove-like bubbling coo or murmur. Black grouse hens visiting the lek decide the overall healthiest male, though not all females may arrive at every lek.
In western Europe, these leks seldom contain more than 40 birds; in Russia, 150 is not uncommon and 200 have been recorded. [14]
When mated successfully, she will fly away from the site to a suitable nesting site with an abundance of dense shrub or tall vegetation, often located at a tree base in between roots, under low branches, beside a boulder, or extremely rarely, a used raptor's or corvid's nest 7 metres (23 ft) off the ground. A dent (23–28 centimetres (9.1–11.0 in) wide by 10–11 centimetres (3.9–4.3 in) deep) is scraped out on the dirt floor and cushioned with grasses, sticks, leaves, and feathers. About 6-11 pale buff eggs speckled brown are then laid in the nest, incubated for approx. 23–28 days. The chicks consume invertebrates, transitioning to more plant matter as they mature. By around 10–14 days and so forth, they are capable of short flights. [13]
Where their range overlaps in similar biomes of other species, they are capable of hybridizing with the ringneck pheasant, western capercaillie, black-billed capercaillie, Siberian grouse, hazel grouse, and willow ptarmigan. [4]
The tails of black-cocks have, since late Victorian times, been popular adornments for hats worn with Highland Dress. Most commonly associated with Glengarry and Balmoral or Tam o' Shanter caps, they still continue to be worn by pipers of civilian and military pipe bands. Since 1904, all ranks of the Royal Scots and King's Own Scottish Borderers have worn them in their full-dress headgear and that tradition is carried on in the dress glengarries of the current Scottish super regiment, the Royal Regiment of Scotland.[ citation needed ]
The dunlin is a small wader, formerly sometimes separated with the other "stints" in the genus Erolia. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from dun, "dull brown", with the suffix -ling, meaning a person or thing with the given quality.
The Eurasian curlew or common curlew is a very large wader in the family Scolopacidae. It is one of the most widespread of the curlews, breeding across temperate Europe and Asia. In Europe, this species is often referred to just as the "curlew", and in Scotland known as the "whaup" in Scots.
The willow ptarmigan is a bird in the grouse subfamily Tetraoninae of the pheasant family Phasianidae. It is also known as the willow grouse and in Ireland and Britain, where the subspecies L. l. scotica was previously considered to be a separate species, as the red grouse. It breeds in birch and other forests and moorlands in northern Europe, the tundra of Scandinavia, Siberia, Alaska and Canada, in particular in the provinces of Newfoundland and Labrador and Quebec. It is the state bird of Alaska. In the summer the birds are largely brown, with dappled plumage, but in the winter they are white with some black feathers in their tails. The species has remained little changed from the bird that roamed the tundra during the Pleistocene. Nesting takes place in the spring when clutches of four to ten eggs are laid in a scrape on the ground. The chicks are precocial and soon leave the nest. While they are young, both parents play a part in caring for them. The chicks eat insects and young plant growth while the adults are completely herbivorous, eating leaves, flowers, buds, seeds and berries during the summer and largely subsisting on the buds and twigs of willow and other dwarf shrubs and trees during the winter.
The western capercaillie, also known as the Eurasian capercaillie, wood grouse, heather cock, cock-of-the-woods, or simply capercaillie, is a heavy member of the grouse family and the largest of all extant grouse species. The heaviest-known specimen, recorded in captivity, had a weight of 7.2 kilograms. Found across Europe and the Palearctic, this primarily-ground-dwelling forest grouse is renowned for its courtship display. The bird shows extreme sexual dimorphism, with males nearly twice the size of females. The global population is listed as "least concern" under the IUCN, although the populations of central Europe are declining and fragmented, or possibly extirpated.
The spruce grouse, also known as Canada grouse, spruce hen or fool hen, is a medium-sized grouse closely associated with the coniferous boreal forests or taiga of North America. It is the only member of the genus Canachites.
The ruffed grouse is a medium-sized grouse occurring in forests from the Appalachian Mountains across Canada to Alaska. It is the most widely distributed game bird in North America. It is non-migratory. It is the only species in the genus Bonasa.
The greater sage-grouse, also known as the sagehen, is the largest grouse in North America. Its range is sagebrush country in the western United States and southern Alberta and Saskatchewan, Canada. It was known as simply the sage grouse until the Gunnison sage-grouse was recognized as a separate species in 2000. The Mono Basin population of sage grouse may also be distinct.
The Gunnison grouse, Gunnison sage-grouse or lesser sage-grouse is a species of grouse endemic to the United States. It is similar to the closely related greater sage-grouse in appearance, but about a third smaller in size, with much thicker plumes behind the head; it also has a less elaborate courtship dance. It is restricted in range to southwestern Colorado and extreme southeastern Utah, with the largest population residing in the Gunnison Basin region in Colorado. Despite being native to a country where the avifauna is relatively well known, it was overlooked until the 1990s due to the similarities with the sage grouse, and only described as a new species in 2000—making it the first new avian species to be described from the USA since the 19th century. The description of C. minimus as a separate species is supported by a molecular study of genetic variation, showing that gene flow between the large-bodied and the small-bodied birds is absent.
The common reed bunting is a passerine bird in the bunting family Emberizidae, a group now separated by most modern authors from the finches, Fringillidae. The genus name Emberiza is from Old German Embritz, a bunting. The specific schoeniclus is from Ancient Greek skhoiniklos, a now unknown waterside bird.
The sharp-tailed grouse, also known as the sharptail or fire grouse, is a medium-sized prairie grouse. One of three species in the genus Tympanuchus, the sharp-tailed grouse is found throughout Alaska, much of Northern and Western Canada, and parts of the Western and Midwestern United States. The sharp-tailed grouse is the provincial bird of the Canadian province of Saskatchewan.
The hazel grouse, sometimes called the hazel hen, is one of the smaller members of the grouse family of birds. It is a sedentary species, breeding across the Palearctic as far east as Hokkaido, and as far west as eastern and central Europe, in dense, damp, mixed coniferous woodland, preferably with some spruce. The bird is sometimes referred to as "rabchick" by early 20th century English speaking travellers to Russia.
The Caucasian grouse or Caucasian black grouse is a large bird in the grouse family. It is closely related to the black grouse.
The Siberian jay is a small jay with a widespread distribution within the coniferous forests in North Eurasia. It has grey-brown plumage with a darker brown crown and a paler throat. It is rusty-red in a panel near the wing-bend, on the undertail coverts and on the sides of the tail. The sexes are similar. Although its habitat is being fragmented, it is a common bird with a very wide range so the International Union for Conservation of Nature has assessed its conservation status as being of "least concern".
The East Siberian taiga ecoregion, in the Taiga and boreal forests biome, is a very large biogeographic region in eastern Russia.
The black-billed capercaillie, also known as eastern capercaillie, Siberian capercaillie, spotted capercaillie or stone capercaillie, is a large grouse species closely related to the more widespread western capercaillie. It is a sedentary species which breeds in the larch taiga forests of eastern Siberia as well as parts of northern Mongolia and China. In the far west of its distribution, the black-billed capercaillie has been known to hybridize with the western capercaillie. Compared to its western cousin, the Siberian capercaillie is also more adaptable to open habitat, given the larch forests it lives in are usually less dense than other taiga communities. Thus, they tend to avoid thick coniferous forests.
Tetrao is a genus of birds in the grouse subfamily known as capercaillies. They are some of the largest living grouse. Feathers from the bird were used to create the characteristic hat of the bersaglieri, an Italian ace infantry formation.
The white-necklaced partridge, also known as the collared partridge or Rickett's hill-partridge, is a species of bird in the family Phasianidae. It is endemic to southeastern China. It is threatened by habitat loss and hunting, and the IUCN has assessed it as near-threatened.
Sage-grouse are grouse belonging to the bird genus Centrocercus. The genus includes two species: the Gunnison grouse and the greater sage-grouse. These birds are distributed throughout large portions of the north-central and Western United States, as well as the Canadian provinces of Alberta and Saskatchewan. The International Union for Conservation of Nature classified the C. minimus species as endangered in 2020 and C. urophasianus as near threatened in 2016.
The Cantabrian capercaillie is a subspecies of the western capercaillie in the grouse family Tetraonidae. It is one of two subspecies found in Spain.
The rackelhahn or rackelwild is a hybrid between the western capercaillie and the black grouse.