Common moorhen

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Common moorhen
Common moorhen (Gallinula chloropus) France.jpg
Adult G. c. chloropus and audio recording of call, both from France
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Gruiformes
Family: Rallidae
Genus: Gallinula
Species:
G. chloropus
Binomial name
Gallinula chloropus
Subspecies

About five; see text

GallinulaChloropusIUCNver2019-2.png
Range of G. chloropus
  Breeding
  Resident
  Non-breeding
  Probably extinct
Synonyms
  • Fulica chloropusLinnaeus, 1758
  • Fulica fuscaLinnaeus, 1766

The common moorhen (Gallinula chloropus), also known as the waterhen or swamp chicken, is a bird species in the rail family (Rallidae). It is distributed across many parts of the Old World, from Africa to Europe and Asia. [1]

Contents

The common moorhen lives around well-vegetated marshes, ponds, canals and other wetlands. [1] The species is not found in the polar regions or many tropical rainforests. Elsewhere it is likely the most common rail species, except for the Eurasian coot in some regions.

The closely related common gallinule of the New World has been recognized as a separate species by most authorities, [2] [ circular reference ] starting with the American Ornithologists' Union and the International Ornithological Committee in 2011. [3]

Taxonomy

The common moorhen was formally described in 1758 by the Swedish naturalist Carl Linnaeus in the tenth edition of his Systema Naturae . He placed it in the genus Fulica and coined the binomial name Fulica chloropus. [4] [5] The common moorhen is now one of five extant species placed in the genus Gallinula that was introduced in 1760 by the French zoologist Mathurin Jacques Brisson. [6] The genus name is from Latin gallinula meaning "little hen" or "little chicken". The specific epithet chloropus combines the Ancient Greek khlōros χλωρός meaning "green" and pous (πούς) meaning "foot". [7]

The name mor-hen has been recorded in English since the 13th century. [8] The word moor here is an old sense meaning marsh; [8] the species is not usually found in moorland. An older name, common waterhen, is more descriptive of the bird's habitat. A "watercock" is not a male "waterhen" but the rail species Gallicrex cinerea , not closely related to the common moorhen. "Water rail" usually refers to Rallus aquaticus , again not closely related.

Five subspecies are recognised: [6]

Description

Common moorhen feet have no webbing. Moorhen feet.jpg
Common moorhen feet have no webbing.

The moorhen is a distinctive species, with predominantly black and brown plumage, with the exception of a white under-tail, white streaks on the flanks, yellow legs and a red frontal shield. The bill is red with a yellow tip. The young are browner and lack the red shield. The frontal shield of the adult has a rounded top and fairly parallel sides; the tailward margin of the red unfeathered area is a smooth waving line. Race meridionalis is smaller than the nominate, has slaty blue-grey upperwing coverts and lacks the olive wash. Race orientalis is similar to meridionalis but has a larger shield. Race pyrrhorrhoa is darker than the nominate; race pyrrhorrhoa has buff undertail coverts . [9] In the related common gallinule (Gallinula galeata) of the Americas, the frontal shield has a fairly straight top and is less wide towards the bill, giving a marked indentation to the back margin of the red area.

The common moorhen gives a wide range of gargling calls and will emit loud hisses when threatened. [10] A midsized to large rail, it can range from 30 to 38 cm (12 to 15 in) in length and span 50 to 62 cm (20 to 24 in) across the wings. The body mass of this species can range from 192 to 500 g (6.8 to 17.6 oz). [11] [12]

Distribution and habitat

This is a common breeding and resident bird in marsh environments, rivers, well-vegetated lakes and even in city parks. Populations in areas where the waters freeze, such as eastern Europe, will migrate to more temperate climates. In China, common moorhen populations are largely resident south of the Yangtze River, whilst northern populations migrate in the winter, therefore these populations show high genetic diversity. [13]

Behaviour

Food and feeding

This species will consume a wide variety of vegetable material and small aquatic creatures. They forage beside or in the water, sometimes walking on lilypads or upending in the water to feed. They are often secretive, but can become tame in some areas. Despite loss of habitat in parts of its range, the common moorhen remains plentiful and widespread.

Breeding

The birds are territorial during breeding season, and will fight with other members of their species, as well as other water birds such as ducks, to drive them out of their territory. The nest is a basket built on the ground in dense vegetation. Laying starts in spring, between mid-March and mid-May in Northern hemisphere temperate regions. About 8 eggs are usually laid per female early in the season; a brood later in the year usually has only 5–8 or fewer eggs. Nests may be re-used by different females. Incubation lasts about three weeks. Both parents incubate and feed the young. These fledge after 40–50 days, become independent usually a few weeks thereafter, and may raise their first brood the next spring. When threatened, the young may cling to the parents' body, after which the adult birds fly away to safety, carrying their offspring with them. [10] [14]

Status and population

Moorhen sighted in Fangu, Corsica (France)

On a global scale – all subspecies taken together – the common moorhen is as abundant as its vernacular name implies. It is therefore considered a species of Least Concern by the IUCN. [1] However, small populations may be prone to extinction. The population of Palau, belonging to the widespread subspecies G. c. orientalis and locally known as debar (a generic term also used for ducks and meaning roughly "waterfowl"), is very rare, and apparently the birds are hunted by locals. Most of the population on the archipelago occurs on Angaur and Peleliu, while the species is probably already gone from Koror. In the Lake Ngardok wetlands of Babeldaob, a few dozen still occur, but the total number of common moorhens on Palau is about in the same region as the Guam population: fewer than 100 adult birds (usually fewer than 50) have been encountered in any survey. [15]

Other localised groups of common moorhen are starting to come under threat. The Royal Society for the Protection of Birds in the United Kingdom has the common moorhen classified as one of its 103 species whose conservation status is of moderate concern [16] due to its recent population decine. The number of breeding pairs has fallen to its lowest level in the UK since 1966 [17] and has been protected under the Wildlife and Countryside Act (1981).

The common moorhen is one of the birds (the other is the Eurasian coot, Fulica atra) from which the cyclocoelid flatworm parasite Cyclocoelum mutabile was first described. [18] The bird is also parasitised by the moorhen flea, Dasypsyllus gallinulae. [19]

Subspecies

Five subspecies are today considered valid; several more have been described that are now considered junior synonyms. Most are not very readily recognizable, as differences are rather subtle and often clinal. Usually, the location of a sighting is the most reliable indication as to subspecies identification, but the migratory tendencies of this species make identifications based on location not completely reliable. In addition to the extant subspecies listed below, an undescribed form from the Early Pleistocene is recorded from Dursunlu in Turkey. [20] [21] [22]

List of subspecies by date of description
Common and
trinomial names
DescriptionRange
Eurasian common moorhen
G. c. chloropus(Linnaeus, 1758)
Includes correiana and indica.
Moorhen 1c (5370646255).jpg
Wings and back blackish-oliveRanges from Northwest Europe to North Africa and eastwards to Central Siberia and from the humid regions of the Indian Subcontinent and Southeast Asia eastwards to Japan; also found the Canary, Azores, Madeira, and Cape Verde islands.
Indo-Pacific common moorhen
G. c. orientalis(Horsfield, 1821)
Small, with slate grey upperwing coverts and large frontal shield.Found in the Seychelles, Andaman Islands, and South Malaysia through Indonesia; also found in the Philippines and Palau. The breeding population existing on Yap in Micronesia since the 1980s is probably of this subspecies, but might be of the rare G. c. guami. [23] [24]
Population size: Perhaps a few 100s on Palau as of the early 2000s, [15] less than 100 on Yap as of the early 2000s. [23] [24]
African common moorhen
G. c. meridionalis(C. L. Brehm, 1831)
Gallinula chloropus meridionalis Marievale 3.jpg
Similar to orientalis, but the frontal shield is smaller.Found in Sub-Saharan Africa and Saint Helena.
Madagascan common moorhen
G. c. pyrrhorrhoa(A. Newton, 1861)
Gallinula chloropus pyrrhorrhoa Mauritius.jpg
Similar to meridionalis, but the undertail coverts are buff.Found on the islands of Madagascar, Réunion, Mauritius, and the Comoros.
Mariana common moorhen
G. c. guami(Hartert, 1917)
Called pulattat in Chamorro.
Body plumage is very dark.Endemic to the Northern Mariana Islands, but see also G. c. orientalis above.
Population size: About 300 as of 2001. [25]

Life cycle

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<span class="mw-page-title-main">Common gallinule</span> Species of bird

The common gallinule is a bird in the family Rallidae. It was split from the common moorhen by the American Ornithologists' Union in July 2011. It lives around well-vegetated marshes, ponds, canals, and other wetlands in the Americas. The common gallinule is one of the most conspicuous rail species in North America, along with the American coot.

<span class="mw-page-title-main">Eurasian coot</span> Species of bird

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<span class="mw-page-title-main">American coot</span> Species of bird

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<span class="mw-page-title-main">Western swamphen</span> Species of waterbird

The western swamphen is a species of swamphen in the rail family Rallidae, one of the six species of purple swamphen. From the French name talève sultane, it is also known as the sultana bird. This chicken-sized bird, with its large feet, bright plumage and red bill and frontal shield is easily recognisable in its native range. It used to be considered the nominate subspecies of the purple swamphen, but is now recognised as a separate species. The western swamphen is found in wetlands in Spain, Portugal, southeastern France, Italy and northwestern Africa.

<span class="mw-page-title-main">American purple gallinule</span> Species of bird

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<span class="mw-page-title-main">Gough moorhen</span> Species of bird

The Gough moorhen is a medium-sized, almost flightless bird that is similar to the common moorhen, but is smaller, stockier, and has shorter wings. The bird has a distinctive yellow-tipped red bill and red frontal shield. Its first account was written in 1888 by the polar explorer George Comer, whom the specific name comeri commemorates. This bird is found only on two remote islands in the South Atlantic.

<span class="mw-page-title-main">Tristan moorhen</span> Extinct species of bird

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<span class="mw-page-title-main">Azure gallinule</span> Species of bird

The azure gallinule is a species of bird in subfamily Rallinae of family Rallidae, the rails, gallinules, and coots. It is found in Argentina, Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Paraguay, Peru, Suriname, Trinidad and Tobago, and Venezuela.

<span class="mw-page-title-main">Hawaiian gallinule</span> Subspecies of bird

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References

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