| Populus tremula | |
|---|---|
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Scientific classification  | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Angiosperms |
| Clade: | Eudicots |
| Clade: | Rosids |
| Order: | Malpighiales |
| Family: | Salicaceae |
| Genus: | Populus |
| Section: | Populus sect. Populus |
| Species: | P. tremula |
| Binomial name | |
| Populus tremula | |
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| Distribution map | |
Populus tremula (commonly called aspen, common aspen, Eurasian aspen, European aspen, or quaking aspen) [2] is a species of poplar native to cool temperate regions of the Old World.
It is a substantial deciduous tree growing to 40 metres (130 ft) tall by 10 m (33 ft) broad, with a trunk attaining over 1 metre (3 ft 3 in) in diameter. [3] The bark is pale greenish-grey and smooth on young trees with dark grey diamond-shaped lenticels, becoming dark grey and fissured on older trees.[ citation needed ]
The adult leaves, produced on branches of mature trees, are nearly round, slightly wider than long, 2–8 cm (1–3 in) diameter, with a coarsely toothed margin and a laterally flattened petiole 4–8 cm (2–3 in) long. The flat petiole allows them to tremble in even slight breezes, and is the source of its scientific name, [3] as well as one of its vernacular names "langues de femmes" attributed to Gerard's 17th-century Herball. The leaves on seedlings and fast-growing stems of suckers (root sprouts) are of a different shape, heart-shaped to nearly triangular. [4] They are also often much larger, up to 20 centimetres (7.9 in) long; their petiole is also less flattened.[ citation needed ]
The flowers are wind-pollinated catkins produced in early spring before the new leaves appear; they are dioecious, with male and female catkins on different trees. The male catkins are patterned green and brown, 5–10 centimetres (2.0–3.9 in) long when shedding pollen; the female catkins are green, 2–6 centimetres (0.79–2.36 in) long at pollination, maturing in early summer to bear 10–20 (50–80) capsules each containing numerous tiny seeds embedded in downy fluff. The fluff assists wind dispersal of the seeds when the capsules split open at maturity. [5] [6] [3]
It can be distinguished from the closely related North American Populus tremuloides, which is nearly identical, by the leaves being more coarsely toothed. [6]
Like other aspens, it spreads extensively by suckers (root sprouts), which may be produced up to 40 m from the parent tree, forming extensive clonal colonies. [5] [6] This often makes the job of clearing unwanted trees from an area especially difficult, as new suckers will continue to sprout from the extensive root system for up to several years after all surface growth has been eliminated.[ citation needed ]
The species is native to Europe and Asia, from Iceland [7] and the British Isles [8] east to Kamchatka, north to inside the Arctic Circle in Scandinavia and northern Russia, and south to central Spain, Turkey, the Tian Shan, North Korea, and northern Japan. It also occurs at one site in northwest Africa in Algeria. In the south of its range, it occurs at high altitudes in mountains. [6] [9]
Eurasian aspen is a water and light demanding species that is able to vigorously colonize an open area after fire, clear cutting or other kind of damages. After an individual has been damaged or destroyed, root suckers are produced abundantly on the shallow lateral roots. Fast growth continues until the age of about 20 years when crown competition increases. After that, growth speed decreases and culminates at about 30 years of age. Aspen can reach an age of 200 years. [3]
It is a very hardy species and tolerates long, cold winters and short summers.[ citation needed ]
Aspen is resistant to browsing pressure by fallow deer due to its unpleasant taste. [10]
This species is important for the hornet moth, which uses it as a host during the larval stage.[ citation needed ]
Fossils of Populus tremula have been described from the fossil flora of Kızılcahamam district in Turkey which is of early Pliocene age. [11]
The aspen is found in cultivation in parks and large gardens. [12] The fastigiate cultivar ‘Erecta’, with bright yellow autumn colouring, has gained the Royal Horticultural Society’s Award of Garden Merit. [13] [14] The cultivar is colloquially known as "Swedish columnar" in Canada and the United States. [15]
The hybrid with Populus alba (white poplar), known as grey poplar, Populus × canescens, is widely found in Europe and central Asia. Hybrids with several other aspens have also been bred at forestry research institutes in order to find trees with greater timber production and disease resistance (e.g. P. tremula × P. tremuloides, bred in Denmark [16] ).
The wood of aspen is light and soft with very little shrinkage. It is used for lumber and matches but is also valued in the pulp and paper industry, being particularly useful for writing paper. In addition, it is used for plywood and different types of flake and particle boards. Given its hardiness and capacity for rapid growth and regeneration, it plays an important role in the production of wood for renewable energy. Ecologically, the species is important as many insect and fungi species benefit from it. The tree further provides habitat for several mammals and birds that require young forests. [3]
Populus is a genus of 25–30 species of deciduous flowering plants in the family Salicaceae, native to most of the Northern Hemisphere. English names variously applied to different species include poplar, aspen, and cottonwood.
Populus section Populus, of the Populus (poplar) genus, includes the aspen trees and the white poplar Populus alba. The five typical aspens are all native to cold regions with cool summers, in the north of the Northern Hemisphere, extending south at high altitudes in the mountains. The White Poplar, by contrast, is native to warmer regions, with hot, dry summers. These trees are all medium-sized deciduous trees ranging 15–30 metres (49–98 ft) tall.
Tilia platyphyllos, the large-leaved lime or large-leaved linden, is a species of flowering plant in the family Malvaceae (Tiliaceae). It is a deciduous tree, native to much of Europe, including locally in southwestern Great Britain, growing on lime-rich soils. The common names largeleaf linden and large-leaved linden are in standard use throughout the English-speaking world except in the British Isles, where it is known as large-leaved lime. The name "lime", possibly a corruption of "line" originally from "lind", has been in use for centuries and also attaches to other species of Tilia. It is not, however, closely related to the lime fruit tree, a species of citrus.
A clonal colony or genet is a group of genetically identical individuals, such as plants, fungi, or bacteria, that have grown in a given location, all originating vegetatively, not sexually, from a single ancestor. In plants, an individual in such a population is referred to as a ramet. In fungi, "individuals" typically refers to the visible fruiting bodies or mushrooms that develop from a common mycelium which, although spread over a large area, is otherwise hidden in the soil. Clonal colonies are common in many plant species. Although many plants reproduce sexually through the production of seed, reproduction occurs by underground stolons or rhizomes in some plants. Above ground, these plants most often appear to be distinct individuals, but underground they remain interconnected and are all clones of the same plant. However, it is not always easy to recognize a clonal colony especially if it spreads underground and is also sexually reproducing.
Populus nigra, the black poplar, is a species of cottonwood poplar, the type species of section Aigeiros of the genus Populus, native to Europe, southwest and central Asia, and northwest Africa.
The balsam poplars are a group of about 10 species of poplars, indigenous to North America and eastern Asia, distinguished by the balsam scent of their buds, the whitish undersides of their leaves, and the leaf petiole being round in cross-section. They are large deciduous trees, 30–60 m tall, with leaves with a rounded base, pointed apex, and a whitish waxy coating on the underside of the leaf; this latter distinguishes them from most other poplars. The name is derived from the pleasant balsam smell of the opening buds and leaves in spring, produced by a sticky gum on the buds which also helps protect the buds from insect damage. The balsam poplars are light-demanding trees that require considerable moisture. Balsam poplars are tolerant of very cold conditions, occurring further north than other poplars except for the aspens. The poplars in Southern California are tolerant of 100 plus degree heat. They grow along dry washes and dry riverbed's. The dry washes and dry riverbeds will have flowing water when it rains sufficiently. Their leaves hang down and are at an edge to the sun. This may be another factor why they can take the high heat. Their leaves tremble in the slightest breeze like the quaking aspen
Populus alba, commonly called silver poplar, silverleaf poplar, or white poplar, is a species of poplar, most closely related to the aspens. It is native to a region spanning from the Atlas Mountains of Africa, through most of South and Central Europe, into Central Asia; it has been introduced to many temperate, moist regions worldwide. It grows in moist sites, often by watersides, in regions with hot summers and cold to mild winters.
Quercus trojana, the Macedonian oak is an oak in the turkey oak section (Quercus sect. Cerris).
Betula populifolia, known as the gray birch, is a deciduous tree in the family Betulaceae. It is native to eastern North America and is most commonly found in the northeast United States as well as southern Quebec, New Brunswick, and Nova Scotia. The tree is a pioneer species that is commonly found in sites following disturbance, such as fire or logging. Gray birches don't have as much economic value as other birch species but are still commonly used as ornamental trees.
Populus tremuloides is a deciduous tree native to cooler areas of North America, one of several species referred to by the common name aspen. It is commonly called quaking aspen, trembling aspen, American aspen, mountain or golden aspen, trembling poplar, white poplar, and popple, as well as others. The trees have tall trunks, up to 25 metres tall, with smooth pale bark, scarred with black. The glossy green leaves, dull beneath, become golden to yellow, rarely red, in autumn. The species often propagates through its roots to form large clonal groves originating from a shared root system. These roots are not rhizomes, as new growth develops from adventitious buds on the parent root system.
Populus trichocarpa, the black cottonwood, western balsam-poplar or California poplar, is a deciduous broadleaf tree species native to western North America. It is used for timber, and is notable as a model organism in plant biology.
Pterocarya fraxinifolia is a species of tree in the Juglandaceae family. It is commonly known as the Caucasian wingnut or Caucasian walnut. It is native to the Caucasian region Armenia, Azerbaijan, Georgia, Iran, Russia, Ukraine and Turkey. It was introduced to France in 1784, and to Great Britain after 1800.
The Crooked Trees, Crooked Bush, Twisted Trees or the Crooked Trees of Alticane are a grove of deformed trembling aspen trees of type Populus tremuloides Michx. found in Saskatchewan, Canada. They are found approximately twenty kilometers north-north-west of the town of Hafford, Saskatchewan and just over five kilometers south-west of Alticane.
Populus grandidentata, commonly called large-tooth aspen, big-tooth aspen, American aspen, Canadian poplar, or white poplar, is a deciduous tree native to eastern North America.
Batrachedra praeangusta is a moth of the family Batrachedridae which is native to Europe. It is also found in North America. It was first described by Adrian Haworth in 1828 from the type specimen found in England. The foodplants of the larvae are poplars and willows.
Epinotia nisella is a moth of the family Tortricidae which is found in the Palearctic, Europe and North America. It was first described by Carl Alexander Clerck in 1759.
Ectoedemia argyropeza is a moth of the family Nepticulidae. It is a widespread species, with a Holarctic distribution.
Aspen is a common name for certain tree species; some, but not all, are classified by botanists in the section Populus, of the Populus genus.
Populus × canescens, the grey poplar, is a hybrid between Populus alba and P. tremula. It is intermediate between its parents, with a thin grey downy coating on the leaves, which are much less deeply lobed than the leaves of P. alba. It is a very vigorous tree with marked hybrid vigour, reaching 40 metres tall and with a trunk diameter over 1.5 m (5 ft) – much larger than either of its parents. Most trees in cultivation are male, but female trees occur naturally and some of these are also propagated.
This fossil flora in Turkey stems from at least six Pliocene deposits in Güvem and Beşkonak villages, 80 kilometres (50 mi) north of Ankara, 22 kilometres (14 mi) north of Kızılcahamam and 125 kilometres (78 mi) south of the Black Sea coast. They are 1,000 metres (3,300 ft) to 1,500 metres (4,900 ft) above sea level, extending 15 kilometres (9.3 mi) north-south and 7 kilometres (4.3 mi) east-west. Between six and seven million years ago, a fresh water lake existed there in a forested area with mostly broad leaved deciduous tree species, dominated by sequoia and oak.
