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Monoplacophora, meaning "bearing one plate", is a polyphyletic superclass of molluscs with a cap-like shell, inhabiting deep sea environments. Extant representatives were not recognized as such until 1952; previously they were known only from the fossil record, and were thought to have become extinct 375 million years ago.
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.
The Coelosclerithophorans are a polyphyletic group of organisms bearing hollow sclerites made of aragonite, and with a supposedly distinctive microstructure.
The evolution of the molluscs is the way in which the Mollusca, one of the largest groups of invertebrate animals, evolved. This phylum includes gastropods, bivalves, scaphopods, cephalopods, and several other groups. The fossil record of mollusks is relatively complete, and they are well represented in most fossil-bearing marine strata. Very early organisms which have dubiously been compared to molluscs include Kimberella and Odontogriphus.
Yochelcionella is an extinct genus of basal molluscs which lived during the Tommotian epoch, the first epoch of the Cambrian period. This genus is often reconstructed to resemble snails.
Helcionellid or Helcionelliformes is an order of small fossil shells that are universally interpreted as molluscs, though no sources spell out why this taxonomic interpretation is preferred. These animals are first found about 540 to 530 million years ago in the late Nemakit-Daldynian age, which is the earliest part of the Cambrian period. A single species persisted to the Early Ordovician. These fossils are component of the small shelly fossils (SSF) assemblages.
Helcionelloida is an extinct group of ancient molluscs. These are the oldest known conchiferan molluscs, that is, they had a mineralised shell. Some members of this class were mistaken for Monoplacophorans. The class was erected by Peel in 1991.
Knightoconus antarcticus is an extinct species of fossil monoplacophoran from the Cambrian Minaret Formation of Antarctica. It is thought to represent an ancestor to the cephalopods. It had a chambered conical shell, but lacked a siphuncle.
Matthevia is a genus of Cambrian molluscs, perhaps related to the chitons. It consists of repeated monoplacophoran-like shells; according to one hypothesis, chitons arose when these tall shells began to overlap over the generations. The tall element of the shell was retained and forms the tips of modern chiton plates. There are distinct head, 'centre', and tail valves, which occur approximately in the ratio 1:5:1 — suggesting a seven-plated configuration.
Micropilina is a genus of monoplacophoran molluscs. They are very small, mostly deepwater animals which have a superficially limpet-like shell.
Khairkhaniidae is an extinct family of Paleozoic fossil molluscs of uncertain position. They are thought to belong to either the Gastropoda or Monoplacophora. They possess planispiral coiled shells with a columnar microstructure.
Yochelcionelloidea is an extinct superfamily of paleozoic molluscs of uncertain position. The earliest yochelcionellids are known from the Middle Tommotian, but they are most diverse from the Botomian through the early Middle Cambrian.
The Kirengellids are a group of problematic Cambrian fossil shells of marine organisms. The shells bear a number of paired muscle scars on the inner surface of the valve.
Anabarella is a species of bilaterally-flattened monoplacophoran mollusc, with a morphological similarity to the rostroconchs. Its shell preserves evidence of three mineralogical textures on its outer surface: it is polygonal near the crest of the shell, subsequently changing to both spiny and stepwise. Its internal microstructure is calcitic and semi-nacreous. Its name reflects its provenance from Anabar, Siberia. It has been interpreted as ancestral to the rostroconchs, and has been aligned to the Helcionellidae.
Fordilla is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early Cambrian fossils found in North America, Greenland, Europe, the Middle East, and Asia. The genus currently contains three described species, Fordilla germanica, Fordilla sibirica, and the type species Fordilla troyensis.
The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.
Stenothecoida is a taxon of bivalved fossils from the Early to middle Cambrian period. They look a bit like brachiopods or bivalve molluscs.
Watsonella is an extinct genus of mollusc known from early (Terreneuvian) Cambrian strata. It has been hypothesized to be close to the origin of bivalves. It contains a single species, Watsonella crosbyi.
Bruce Norman Runnegar is an Australian-born paleontologist and professor at UCLA. His research centers on using the fossil record to determine how, where, and when life originated and evolved. He has published on a wide variety of topics, including the phylogeny of molluscs, Dickinsonia fossils and oxygen levels, and molecular clock techniques.
Protoconodonts are an extinct group of Cambrian animals known from fossilized phosphatic tooth-like structures. They were originally described as an informal group of early conodonts, though more recent studies consider them to be more closely related to chaetognaths.
References
- 1 2 3 Yochelson, E. L. (1969). "Stenothecoida, A Proposed New Class of Cambrian Mollusca". Lethaia. 2: 49–62. doi:10.1111/j.1502-3931.1969.tb01250.x.
- ↑ Runegar & Jell. 1980. Alcheringa, 4(2): 111.
- 1 2 Runnegar, B.; Pojeta Jr, J. (Oct 1974). "Molluscan Phylogeny: the Paleontological Viewpoint". Science. 186 (4161): 311–317. Bibcode:1974Sci...186..311R. doi:10.1126/science.186.4161.311. JSTOR 1739764. PMID 17839855.
- ↑ Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia . 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
- 1 2 Parkhaev P. Yu. 2001. Molluscs and siphonoconchs. In: Alexander E. M. et al. (eds.) The Cambrian biostratigraphy of the Stansbury basin, South Australia. Transactions of the Paleontological Institute, Russian Academy of Sciences, 282: 133–210, plates 24–54. Watsonellinae on the page 187.
- ↑ Budd, G. E.; Jensen, S. (2000). "A critical reappraisal of the fossil record of the bilaterian phyla". Biological Reviews of the Cambridge Philosophical Society. 75 (2): 253–95. doi:10.1111/j.1469-185X.1999.tb00046.x. PMID 10881389.
- ↑ (in Russian) Missarzhevsky. (after 10 July) 1989. Drevneishie skeletnye okamenelosti i stratigrafiia pogranichnykh tolshch Dokembriia i Kembiia. (English translation: Oldest skeletal fossils and stratigraphy of Precambrian and Cambrian boundary beds.) Trudy Geologicheskogo Instituta, Akademia Nauk SSSR, 443, 237 pp., 32 plates. Mellopegmidae is on the page 179.
- ↑ Grabau A. W. 1900. Palaeontology of the Cambrian terranes of the Boston Basin. Occasional Papers of the Boston Society of Natural History 4:601–694.
