| Trimerorhachidae | |
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| Skull of Trimerorhachis insignis | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Order: | † Temnospondyli |
| Suborder: | † Dvinosauria |
| Family: | † Trimerorhachidae Cope, 1882 |
Trimerorhachidae is a family of dvinosaurian temnospondyls, including Lafonius , Trimerorhachis , Procuhy and Neldasaurus . [1] [2] [3]
They were semiaquatic carnivores that lived from the Late Carboniferous to the Early Permian. [4]
Dissorophidae is an extinct family of medium-sized temnospondyls that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America.
Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and armour-like bony plates that distinguish them from the modern soft-bodied lissamphibians.
Dvinosaurs are one of several new clades of temnospondyls named in the phylogenetic review of the group by Yates and Warren 2000. They represent a group of primitive semi-aquatic to completely aquatic temnospondyls, and are known from the Late Carboniferous to the Early Triassic, being most common in the Permian period. Their distinguishing characteristics are a reduction of the otic notch; the loss of a flange on the rear side of the pterygoid; and 28 or more presacral vertebrae.
The Stereospondyli are a group of extinct temnospondyl amphibians that existed primarily during the Mesozoic period. They are known from all seven continents and were common components of many Triassic ecosystems, likely filling a similar ecological niche to modern crocodilians prior to the diversification of pseudosuchian archosaurs.
Dvinosaurus is an extinct genus of amphibious temnospondyls localized to regions of western and central Russia during the middle and late Permian, approximately 265-254 million years ago. Its discovery was first noted in 1921 by Russian paleontologist Vladimir Prokhorovich Amalitskii in a posthumously published paper that documents the findings of a site in Russia's Arkhangelsk District. Its name is derived from the proximity of this site to the Northern Dvina River.
Australerpeton is an extinct genus of stereospondylomorph temnospondyl currently believed to belong to the family Rhinesuchidae. When first named in 1998, the genus was placed within the new family Australerpetontidae. However, studies published a few years later questioned the systematics used in the original description and included the genus within Archegosauridae. A study by Dias & Schultz (2003) reassigned Australerpeton to the family Rhinesuchidae within the suborder Stereospondyli based on an earlier evaluation of the family. In this study, the close similarities between Australerpeton and archegosaurids were attributed to convergent evolution as a result of similar semi-aquatic lifestyles. A redescription of the skeleton of this genus was published by Eltink & Langer in 2014, and the skull was redescribed in a follow-up study published by Eltink et al. in 2016. These studies, as well as a 2017 study focusing on rhinesuchids in general, confirmed that Australerpeton was a rhinesuchid rather than an archegosaurid. Fossils of the genus have been found in the Rio do Rasto Formation of Brazil.
Trematopidae is a family of dissorophoid temnospondyls spanning the late Carboniferous to the early Permian. Together with Dissorophidae, the family forms Olsoniformes, a clade comprising the medium-large terrestrial dissorophoids. Trematopids are known from numerous localities in North America, primarily in New Mexico, Oklahoma, and Texas, and from the Bromacker quarry in Germany.
Limnarchia is a clade of temnospondyls. It includes the mostly Carboniferous-Permian age Dvinosauria and the mostly Permian-Triassic age Stereospondylomorpha. The clade was named in a 2000 phylogenetic analysis of stereospondyls and their relatives. Limnarchia means "lake rulers" in Greek, in reference to their aquatic lifestyles and long existence over a span of approximately 200 million years from the Late Carboniferous to the Early Cretaceous. In phylogenetic terms, Limnarchia is a stem-based taxon including all temnospondyls more closely related to Parotosuchus than to Eryops. It is the sister group of the clade Euskelia, which is all temnospondyls more closely related to Eryops than to Parotosuchus. Limnarchians represent an evolutionary radiation of temnospondyls into aquatic environments, while euskelians represent a radiation into terrestrial environments. While many euskelians were adapted to life on land with strong limbs and bony scutes, most limnarchians were better adapted for the water with poorly developed limbs and lateral line sensory systems in their skulls.
Acroplous is an extinct genus of dvinosaurian Temnospondyli within the family Eobrachyopidae.
Neldasaurus is an extinct genus of dvinosaurian temnospondyl within the family Trimerorhachidae.
Trimerorhachis is an extinct genus of dvinosaurian temnospondyl within the family Trimerorhachidae. It is known from the Early Permian of the southwestern United States, with most fossil specimens having been found in the Texas Red Beds. The type species of Trimerorhachis, T. insignis, was named by American paleontologist Edward Drinker Cope in 1878. Cope named a second species from Texas, T. mesops, in 1896. The species T. rogersi and T. greggi are also from Texas, and the species T. sandovalensis is from New Mexico.
Slaugenhopia is an extinct genus of dvinosaurian temnospondyl within the family Tupilakosauridae. Fossils have been found from the Early Permian San Angelo Formation in Texas. The type and only species, S. texensis, was named in 1962. It may be closely related to the dvinosaur Kourerpeton. Slaugenhopia was once classified as a trimerorhachid but is now classified as a tupilakosaurid.
Olsoniformes is an extinct clade of dissorophoid temnospondyls, including the families Dissorophidae and Trematopidae. Most members of the clade were highly adapted to a terrestrial lifestyle. The clade was named in 2008 and is defined as the least inclusive clade containing Dissorophus multicinctus and Acheloma cumminsi but not Amphibamus grandiceps, Micromelerpeton credneri, or Apateon pedestris. Olsoniforms share various features such as a stout and low ilium and a thin cultriform process. The earliest-branching olsoniform is Palodromeus bairdi, from the Late Carboniferous of Ohio.
Parioxys is an extinct genus of temnospondyl amphibian from the Early Permian of Texas.
The Waggoner Ranch Formation is a geologic formation in northern Texas. It preserves fossils dating back to the Artinskian to Kungurian stages of the Permian period.
The Archer City Formation is a geological formation in north-central Texas, preserving fossils from the Asselian and early Sakmarian stages of the Permian period. It is the earliest component of the Texas red beds, introducing an tropical ecosystem which will persist in the area through the rest of the Early Permian. The Archer City Formation is preceded by the cool Carboniferous swamp sediments of the Markley Formation, and succeeded by the equally fossiliferous red beds of the Nocona Formation. The Archer City Formation was not named as a unique geological unit until the late 1980s. Older studies generally labelled its outcrops as the Moran or Putnam formations, which are age-equivalent marine units to the southwest.
The Arroyo Formation, sometimes termed the Lower Clear Fork Formation, is a geologic formation in Texas. It preserves fossils dating back to the Kungurian stage of the Permian period. It is the lower-most portion of the Clear Fork Group, part of a series of fossiliferous Permian strata in the south-central United States known as the red beds.
Amphibamiformes is an unranked clade with Dissorophoidea created by Schoch (2018). It encompasses all of the taxa traditionally considered to be "amphibamids", branchiosaurids, and hypothetically lissamphibians under the traditional temnospondyl hypothesis of lissamphibian origins. These taxa are typically small-bodied dissorophoids and form the sister group to Olsoniformes, which comprises dissorophids and trematopids.
Richards Spur is a Permian fossil locality located at the Dolese Brothers Limestone Quarry north of Lawton, Oklahoma. The locality preserves clay and mudstone fissure fills of a karst system eroded out of Ordovician limestone and dolomite, with the infilling dating to the Artinskian stage of the early Permian (Cisuralian), around 289 to 286 million years ago. Fossils of terrestrial animals are abundant and well-preserved, representing one of the most diverse Paleozoic tetrapod communities known. A common historical name for the site is Fort Sill, in reference to the nearby military base. Fossils were first reported at the quarry by workers in 1932, spurring a wave of collecting by local and international geologists. Early taxa of interest included the abundant reptile Captorhinus and microsaurs such as Cardiocephalus and Euryodus. Later notable discoveries include Doleserpeton, the most diverse assortment of parareptiles in the Early Permian, and the rare early diapsid Orovenator.
Procuhy is an extinct genus of dvinosaurian temnospondyl amphibian in the family Trimerorhachidae represented by the type species Procuhy nazariensis from the Early Permian of Brazil.
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