Uralosuchus Temporal range: Late Permian | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Temnospondyli |
Family: | † Archegosauridae |
Subfamily: | † Melosaurinae |
Genus: | † Uralosuchus Gubin, 1993 |
Type species | |
†Uralosuchus tverdochlebovae Gubin, 1993 |
Uralosuchus is an extinct genus of temnospondyl amphibian from the Late Permian of Russia, belonging to the group Archegosauroidea. It is a member of the archegosauroidean subfamily Melosaurinae. Fossils have been found in Orenburg Oblast. Uralosuchus was named in 1993 with the description of the type species U. tverdochlebovae. [1]
Below is a cladogram modified from Ruta et al. (2007) showing the relationship of Uralosuchus to other archegosauroids: [2]
Archegosauroidea |
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Lepospondyli is a diverse taxon of amphibian tetrapods. With the exception of one late-surviving lepospondyl from the Late Permian of Morocco, lepospondyls lived from the Early Carboniferous (Mississippian) to the Early Permian and were geographically restricted to what is now Europe and North America. Five major groups of lepospondyls are known: Adelospondyli; Aïstopoda; Lysorophia; Microsauria; and Nectridea. Lepospondyls have a diverse range of body forms and include species with newt-like, eel- or snake-like, and lizard-like forms. Various species were aquatic, semiaquatic, or terrestrial. None were large, and they are assumed to have lived in specialized ecological niches not taken by the more numerous temnospondyl amphibians that coexisted with them in the Paleozoic. Lepospondyli was named in 1888 by Karl Alfred von Zittel, who coined the name to include some tetrapods from the Paleozoic, that shared some specific characteristics in the notochord and teeth. Lepospondyls have sometimes been considered to be either related or ancestral to modern amphibians or to Amniota.
Peltobatrachus is an extinct genus of temnospondyl amphibian from the late Permian period of Tanzania.
Otsheria is an extinct genus of anomodont from the Permian of Russia. Otsheria had a 10 centimetres (3.9 in) long skull.
Edops is an extinct genus of temnospondyl amphibian from the Early Permian Period. Unlike more advanced temnospondyls of the time, such as Eryops, Edops exhibited an archaic pattern of palatal bones, and still possessed various additional bones at the back of the skull. Edopoids also had particularly big premaxillae and proportionally small external nostrils. Within the clade, the most basal member seems to be Edops from the Early Permian Archer City Formation of the US, a broad-skulled animal with large palatal teeth.
Archegosauridae is a family of relatively large and long snouted temnospondyls that lived in the Permian period. They were fully aquatic animals, and were metabolically and physiologically more similar to fish than modern amphibians.
Intasuchus is an extinct genus of temnospondyl amphibian from the Late Permian of Russia. It is known from a single species, Intasuchus silvicola, which was named in 1956. Intasuchus belongs to the family Intasuchidae and is probably its sole member, although other taxa such as Syndyodosuchus and Cheliderpeton have been assigned to the family in the past. Intasuchus most likely belongs to the group Archegosauroidea, Permian relatives of the large, mostly Mesozoic temnospondyl clade Stereospondyli.
Archosaurus is an extinct genus of carnivorous archosauriform reptile. From the latest Permian of Russia and Poland, it is one of the earliest known archosauriforms. The type and only species is Archosaurus rossicus, known from several fragmentary specimens which cumulatively represent parts of the skull and cervical vertebrae.
Konzhukovia is an amphibian genus that belongs to an extinct group of temnospondyls, the largest clade of basal tetrapods including about 198 genera, 292 species, and more than half of which were alive during the early Mesozoic period. The animal was a predator that lived about 260 million years ago, and could get up to about 3 meters in length. Specifically, Konzukovia lived during the Permian, between 252 and 270 million years ago according to the type of rock the fossil was found in. There are three species within this genus, K. vetusta, K. tarda, and K. sangabrielensis, the first two originating from Russia while the latest originating from Southern Brazil. The discovery of this specimen in Southern Brazil provided more evidence to support the idea that during this animals existence, there was a “biological corridor” because of the supercontinent Pangea, allowing these species to be found so far apart from each other. Konzhukovia belongs to the family Archegosauridae, a family consisted of large temnospondyls that most likely compare to modern day crocodiles. Since the discovery of the latest species, K. sangabrielensis Pacheco proposes that there must be the creation of a new family, Konzhokoviidae, a monophyletic group in a sister-group relationship with Stereospondlyi in order to accommodate the three species. Konzhukovia skulls usually exhibit typical rhinesuchid features including an overall parabolic shape, small orbits located more posteriorly, and the pterygoids do not reach the vomer. These animals were long-snouted amphibians that had clear adaptations made for fish catching, as well as exemplifying aquatic features.
Brachyopoidea is a superfamily of temnospondyls that lived during the Mesozoic. It contains the families Brachyopidae and Chigutisauridae. The earliest records of brachyopids are from the Lower Triassic in Australia. The latest-surviving member of the superfamily is the chigutisaurid Koolasuchus from the Early Cretaceous of Australia.
Limnarchia is a clade of temnospondyls. It includes the mostly Carboniferous-Permian age Dvinosauria and the mostly Permian-Triassic age Stereospondylomorpha. The clade was named in a 2000 phylogenetic analysis of stereospondyls and their relatives. Limnarchia means "lake rulers" in Greek, in reference to their aquatic lifestyles and long existence over a span of approximately 200 million years from the Late Carboniferous to the Early Cretaceous. In phylogenetic terms, Limnarchia is a stem-based taxon including all temnospondyls more closely related to Parotosuchus than to Eryops. It is the sister group of the clade Euskelia, which is all temnospondyls more closely related to Eryops than to Parotosuchus. Limnarchians represent an evolutionary radiation of temnospondyls into aquatic environments, while euskelians represent a radiation into terrestrial environments. While many euskelians were adapted to life on land with strong limbs and bony scutes, most limnarchians were better adapted for the water with poorly developed limbs and lateral line sensory systems in their skulls.
Bukobaja is an extinct genus of mastodonsaurid temnospondyl from the middle Triassic of Russia. It contains a single species, Bukobaja enigmatica. Bukobaja mainly occurs in the Bukobay Svita as part of the Ladinian?-age "Mastodonsaurus fauna", a section of Russian Triassic biostratigraphy characterized by "Mastodonsaurus" torvus. It was also present in the underlying Donguz Svita. Bukobaja appears to be a valid genus similar to, yet distinct from, Mastodonsaurus. Despite appearing to possess several unique features, Bukobaja is still known from very few remains. This has led to difficulties in determining its relations more precisely than "Mastodonsauridae incertae sedis". It has also been compared to trematosaurids.
Gobiops is an extinct genus of temnospondyl from the Jurassic of Mongolia, China, and possibly Kyrgyzstan. The genus is represented by a single species, Gobiops desertus. It was named in 1991 from the Late Jurassic Shar Teeg Beds of Mongolia. Additional material was described in 2005 from the Middle Jurassic Toutunhe Formation in the Junggar Basin of China. Gobiops belongs to the family Brachyopidae. The poorly known genus Ferganobatrachus, named in 1990 from Shar Teeg, is probably synonymous with Gobiops.
Thoosuchus is an extinct genus of basal trematosauroid trematosaurian temnospondyl. Fossils have been found from Russia and date back to the Early Triassic. It is the type genus of the family Thoosuchidae, formerly called the subfamily Thoosuchinae and placed within Benthosuchidae. The benthosuchids were originally composed of the majority of basal trematosaurian forms regarded as the ancestors of the trematosaurids. Although the genus was first named in 1940, material from one species, E. yakovlevi, was originally tentatively referred to Trematosuchus in 1926.
Vjushkovisaurus is an extinct genus of Middle Triassic archosauriform. It is known from the Anisian-aged Donguz Gorizont in Sol-Iletsk, Orenburg Oblast, Russia. The genus was named in 1982, with the type species being V. berdjanensis. Material has been collected in the Berdyanka II locality from a fossil assemblage called the Eryosuchus Fauna along the Berdyanka River, specifically in a sand-carbonate concretion in the upper part of the main river channel. Vjushkovisaurus is known only from the holotype PIN 2865/62, a partial postcranial skeleton which consists of 12 presacral vertebrae, left humerus, ribs, a fragment of the coracoid and a fragment of the fibula.
Rhytidostea is a clade of stereospondyl temnospondyls. It was erected in 2000 to include several temnospondyl groups distinct from the "higher" group of capitosaurs, including lydekkerinids, brachyopoids, and rhytidosteids. Rhytidosteans first appeared in the Permian period and underwent an evolutionary radiation during the Induan stage of the Early Triassic. Along with capitosaurs, rhytidosteans comprise much of the larger suborder Stereospondyli. Rhytidostea has often been considered the sister group of the clade Capitosauria, but has been placed in various other phylogenetic positions. In many studies, members of Rhytidostea are split, with lydekkerinids having a more basal position among stereospondyls while rhytidosteids and brachyopoids form a group placed among the more derived trematosaurian stereospondyls.
Vonhuenia is an extinct genus of basal archosauriform from the Early Triassic of Russia. Fossils have been found in the Vokhminskaya Formation, along the Vetluga River that are Induan in age, making Vonhuenia one of the earliest archosauriforms.
Melosaurinae is an extinct subfamily of temnospondyl amphibians, part of the family Archegosauridae. Most melosaurines are known from Russia, although an unnamed species has been found in Brazil. Unlike the long-skulled archegosaurids of the family Platyoposaurinae, melosaurines have shorter, broadened snouts.
Koinia is an extinct genus of temnospondyl amphibian from the Late Permian of Russia. It is an archegosauroid in the subfamily Melosaurinae. Koinia was named in 1993 with the description of the type species K. silantjevi, based on fossils that were found in the Ocher Assemblage Zone, near the Vym River in the Komi Republic.
Kashmirosaurus is an extinct genus of temnospondyl amphibian known from Permo-Carboniferous deposits in the region of Kashmir. It was originally named by English paleontologist Arthur Smith Woodward in 1905 as a species of Archegosaurus called Archegosaurus ornatus. More recently, the species has been recognized as being distinct from Archegosaurus, and it was placed in its own genus Kashmirosaurus in 1996. An additional species of Archegosaurus, A. kashmiriensis, was named in 1960 from the same deposits in Kashmir, and is now considered synonymous with Kashmirosaurus ornatus.
Phthinosuchia is an extinct group of therapsids including two poorly known species, Phthinosuchus discors and Phthinosaurus borrisiaki, from the Middle Permian of Russia. Phthinthosuchus is known a partial crushed skull and Phthinosaurus is known from an isolated lower jaw. The two species have traditionally been grouped together based on their shared primitive characteristics, but more recent studies have proposed that they are more distantly related. Phthinosuchus is either a carnivorous gorgonopsian relative or an anteosaurian dinocephalian while Phthinosaurus is either a herbivorous rhopalodont dinocephalian or a therocephalian.
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