Trimma

Trimma
Trimma gigantum
Trimma caesiura
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Gobiiformes
Family:	Gobiidae
Genus:	Trimma D. S. Jordan & Seale, 1906
Type species
Trimma caesiura D. S. Jordan & Seale, 1906

Trimma is a genus of fish in the family Gobiidae native to the Indian and Pacific Ocean. Together with members of the genus Eviota , they are known commonly as pygmygobies or dwarfgobies.

Species

Trimma caudomaculatum

Trimma habrum

Trimma lantana

Trimma nasa

Trimma naudei female 23.3 mm, Nha Trang, Vietnam

Trimma okinawae

Trimma tevegae

There are currently 111 recognized species in this genus: ^[1]

• Trimma abyssum G. R. Allen, 2015 (Deep-reef pygmygoby) ^[2]
• Trimma agrena R. Winterbottom & I. S. Chen, 2004 (Fishnet pygmygoby)
• Trimma anaima R. Winterbottom, 2000 (Sharp-eye pygmygoby) ^[3]
• Trimma annosum R. Winterbottom, 2003 (Grey-bearded pygmygoby) ^[3]
• Trimma anthrenum R. Winterbottom, 2006 (Honey-bee pygmygoby)
• Trimma aturirii R. Winterbottom, Erdmann & Cahyani, 2015 (Aturiri's pygmygoby) ^[4]
• Trimma avidori (Goren, 1978) (Avidor's pygmygoby)
• Trimma barralli R. Winterbottom, 1995 (Barrall's pygmygoby)
• Trimma bathum R. Winterbottom, 2017 (Deep-reef pygmygoby)
• Trimma benjamini R. Winterbottom, 1996 (Ring-eye pygmygoby) ^[3]
• Trimma bisella R. Winterbottom, 2000 (Two-saddle pygmygoby)
• Trimma blematium R. Winterbottom & Erdmann, 2018 (Blue-eyed pygmygoby)
• Trimma burridgeae R. Winterbottom, 2016 ^[5] (Mary's pygmygoby)
• Trimma caesiura D. S. Jordan & Seale, 1906 (Caesiura pygmygoby) ^[3]
• Trimma cana R. Winterbottom, 2004 (Candycane pygmygoby)
• Trimma capostriatum (Goren, 1981) (Spotted red-lined pygmygoby) ^[3]
• Trimma caudipunctatum T. Suzuki & Senou, 2009 (Spot-tailed pygmygoby)
• Trimma caudomaculatum Yoshino & Araga, 1975 ^[5] (Blotch-tailed pygmygoby)
• Trimma cavicapum R. Winterbottom, Eleanor Brighton & Christophe Mason-Parker, 2024 ^[6]
• Trimma cheni R. Winterbottom, 2011 (Face-stripe pygmygoby)
• Trimma chledophilum G. R. Allen, 2015 (Mud pygmygoby) ^[2]
• Trimma citrum R. Winterbottom & Pyle, 2022 (Lemon pygmygoby) ^[7]
• Trimma corallinum (J. L. B. Smith, 1959) (Polka-dot pygmygoby)
• Trimma corerefum R. Winterbottom, 2016 ^[5] (Colin's pygmygoby)
• Trimma dalerocheila R. Winterbottom, 1984 (Hotlips pygmygoby)
• Trimma emeryi R. Winterbottom, 1985 (Emery's pygmygoby) ^[3]
• Trimma erdmanni R. Winterbottom, 2011 (Erdmann's pygmygoby)
• Trimma erwani Viviani, J. T. Williams & Planes, 2016 (Slanted pygmygoby) ^[8]
• Trimma fangi R. Winterbottom & I. S. Chen, 2004 (Fang's pygmygoby) ^[3]
• Trimma fasciatum T. Suzuki, Sakaue & Senou, 2012 ^[9] (Yellow-banded pygmygoby)
• Trimma filamentosum R. Winterbottom, 1995 (Long-spine pygmygoby)
• Trimma finistrinum R. Winterbottom, 2017 (Porthole pygmygoby)
• Trimma fishelsoni Goren, 1985 (Fishelson's pygmygoby)
• Trimma flammeum (J. L. B. Smith, 1959) (Flame pygmygoby)
• Trimma flavatrum K. Hagiwara & R. Winterbottom, 2007 (Wasp pygmygoby) ^[3]
• Trimma flavicaudatum (Goren, 1982) (Yellow-tailed pygmygoby)
• Trimma fraena R. Winterbottom, 1984 (Bridle pygmygoby)
• Trimma fucatum R. Winterbottom & Southcott, 2007 (Harlot pygmygoby)
• Trimma gigantum R. Winterbottom & Zur, 2007 (Giant pygmygoby)
• Trimma grammistes (Tomiyama, 1936) (Black-striped pygmygoby)
• Trimma griffithsi R. Winterbottom, 1984 (Griffiths' pygmygoby)
• Trimma habrum R. Winterbottom, 2011 (Delicate pygmygoby)
• Trimma haima R. Winterbottom, 1984 (Cut-face pygmygoby)
• Trimma haimassum R. Winterbottom, 2011 (Blood-spot pygmygoby)
• Trimma halonevum R. Winterbottom, 2000 (Pimple pygmygoby)
• Trimma hamartium R. Winterbottom, 2018 (Mistaken pygmygoby)
• Trimma hayashii K. Hagiwara & R. Winterbottom, 2007 (Four-eye pygmygoby)
• Trimma helenae R. Winterbottom, Erdmann & Cahyani, 2014 (Helen's pygmygoby) ^[10]
• Trimma hoesei R. Winterbottom, 1984 (Fork-tail pygmygoby) ^[3]
• Trimma hollemani R. Winterbottom, 2016 ^[5] (Holleman’s pygmygoby)
• Trimma hotsarihiensis R. Winterbottom, 2009 (Helen Reef pygmygoby)
• Trimma imaii T. Suzuki & Senou, 2009 (Imai's pygmygoby)
• Trimma insularum R. Winterbottom & Hoese, 2015 (Cocos pygmygoby) ^[3]
• Trimma irinae R. Winterbottom, 2014 (Irina's pygmygoby) ^[11]
• Trimma kardium R. Winterbottom, Erdmann & Cahyani, 2015 (Heart pygmygoby) ^[4]
• Trimma kitrinum R. Winterbottom & Hoese, 2015 (Citron pygmygoby) ^[3]
• Trimma kudoi T. Suzuki & Senou, 2008 (Kudo’s pygmygoby)
• Trimma lantana R. Winterbottom & C. A. Villa, 2003 (Lantana pygmygoby) ^[3]
• Trimma longispinum R. Winterbottom, 2023 (Long-spined pygmygoby)
• Trimma luteum Viviani, J. T. Williams & Planes, 2016 (Yellow-barred pygmygoby) ^[8]
• Trimma macrophthalmus (Tomiyama, 1936) (Large-eye pygmygoby) ^[3]
• Trimma maiandros Hoese, R. Winterbottom & Reader, 2011 (Zigzag pygmygoby) ^[3]
• Trimma marinae R. Winterbottom, 2005 (Princess pygmygoby)
• Trimma matsunoi T. Suzuki, Sakaue & Senou, 2012 ^[9] (Matsuno's pygmygoby)
• Trimma meityae R. Winterbottom & Erdmann, 2018 (Meity’s pygmygoby)
• Trimma mendelssohni (Goren, 1978) (Mendelssohn's pygmygoby)
• Trimma meranyx R. Winterbottom, Erdmann & Cahyani, 2014 (Day-night pygmygoby) ^[12]
• Trimma meristum R. Winterbottom & Hoese, 2015 (Split-ray pygmygoby) ^[3]
• Trimma milta R. Winterbottom, 2002 (Red-earth pygmygoby) ^[3]
• Trimma multiclitellum G. R. Allen, 2015 (Multi-saddle pygmygoby) ^[2]
• Trimma nasa R. Winterbottom, 2005 (Nasal-bar pygmygoby) ^[3]
• Trimma nauagium G. R. Allen, 2015 (Shipwreck pygmygoby) ^[2]
• Trimma naudei J. L. B. Smith, 1957 (Naude's pygmygoby)
• Trimma necopinum (Whitley, 1959) (Australian pygmygoby) ^[3]
• Trimma nomurai T. Suzuki & Senou, 2007 (Lilac pygmygoby) ^[3]
• Trimma okinawae (Aoyagi, 1949) (Orange-red pygmygoby) ^[3]
• Trimma omanense R. Winterbottom, 2000 (Crescent-wing pygmygoby)
• Trimma pajama R. Winterbottom, Erdmann & Cahyani, 2014 (Pajama pygmygoby) ^[12]
• Trimma panemorfum R. Winterbottom & Pyle, 2004 ^[13] (Sublime pygmygoby)
• Trimma papayum R. Winterbottom, 2011 (Pawpaw pygmygoby)
• Trimma pentherum R. Winterbottom & Hoese, 2015 (Mourning pygmygoby) ^[3]
• Trimma preclarum R. Winterbottom, 2006 (Exquisite pygmygoby) ^[3]
• Trimma quadrimaculatum Hoese, Bogorodsky & A. O. Mal, 2015 (Four-spotted pygmygoby) ^[14]
• Trimma randalli R. Winterbottom & Zur, 2007 (Randall's pygmygoby)
• Trimma readerae R. Winterbottom & Hoese, 2015 ^[3] (Reader's pygmygoby)
• Trimma rubromaculatum G. R. Allen & Munday, 1995 (Red-spotted dwarfgoby)
• Trimma sanguinellus R. Winterbottom & Southcott, 2007 (Sanguinello pygmygoby)
• Trimma sheppardi R. Winterbottom, 1984 (Sheppard's pygmygoby)
• Trimma sostra R. Winterbottom, 2004 (Sostra's pygmygoby)
• Trimma squamicana R. Winterbottom, 2004 (Red-blotch pygmygoby)
• Trimma stobbsi R. Winterbottom, 2001 (Stobb's pygmygoby) ^[3]
• Trimma striatum (Herre, 1945) (Red-lined pygmygoby) ^[3]
• Trimma tauroculum R. Winterbottom & Zur, 2007 (Bulls-eye pygmygoby)
• Trimma taylori Lobel, 1979 (Yellow pygmygoby) ^[3]
• Trimma tevegae Cohen & W. P. Davis, 1969 (Blue-stripe pygmygoby) ^[5]
• Trimma trioculatum R. Winterbottom, Erdmann & Cahyani, 2015 (Three-eyed pygmygoby) ^[4]
• Trimma ukkriti R. Winterbottom, 2021 (Ukkrit’s pygmygoby)
• Trimma unisquame (Gosline, 1959) (Black-margin pygmygoby) ^[3]
• Trimma volcana R. Winterbottom, 2003 (Volcano pygmygoby)
• Trimma wangunui R. Winterbottom & Erdmann, 2019 (Wangunu's pygmygoby)
• Trimma winchi R. Winterbottom, 1984 (Winch's pygmygoby)
• Trimma winterbottomi J. E. Randall & Downing, 1994 (Winterbottom's pygmygoby)
• Trimma woutsi R. Winterbottom, 2002 (Wouter's pygmygoby)
• Trimma xanthochrum R. Winterbottom, 2011 (Step-spot pygmygoby)
• Trimma xanthum R. Winterbottom & Hoese, 2015 (Yellow-red pygmygoby) ^[3]
• Trimma yanagitai T. Suzuki & Senou, 2007 (Brown pygmygoby)
• Trimma yanoi T. Suzuki & Senou, 2008 (Pudgy pygmygoby)
• Trimma yoshinoi T. Suzuki, Ko. Yano & Senou, 2015 ^[15] (Yoshino's pygmygoby)
• Trimma zurae R. Winterbottom, Erdmann & Cahyani, 2014 (Marg's pygmygoby) ^[12]

References

1. Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Trimma". FishBase . October 2024 version.
2. Allen, G.R. (2015). "Four new species of Trimma (Pisces: Gobiidae) from Papua New Guinea and Indonesia" (PDF). Journal of the Ocean Science Foundation. 14: 28–48.
3. Winterbottom, R.; Hoese, D.F. (2015). "A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species" (PDF). Zootaxa. 3934 (1): 1–102. doi:10.11646/zootaxa.3934.1.1. PMID   25781853.
4. Winterbottom, R.; Erdmann, M.V.; Dita Cahyani, N.K. (2015). "New species of Trimma (Actinopterygii, Gobiidae) from Indonesia, with comments on head papillae nomenclature" (PDF). Zootaxa. 3973 (2): 201–226. doi:10.11646/zootaxa.3973.2.1. PMID   26249856.
5. Winterbottom, R. (2016). "Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific". Zootaxa. 4144 (1): 1–53. doi:10.11646/zootaxa.4144.1.1. PMID   27470835.
6. Winterbottom, R., Brighton, E. & Mason-Parker, C. (2024) A new species of Trimma (Teleostei: Gobiidae) from the Seychelles, Indian Ocean. Journal of the Ocean Science Foundation, 41, 37-45.https://doi.org/10.5281/zenodo.12594852
7. Winterbottom, R. & Pyle, R.L. (2022) A new species of Trimma (Teleostei: Gobiidae) from the deep reefs of Vanuatu, western Pacific Ocean. Journal of the Ocean Science Foundation, 39, 2-8.; https://doi.org/10.5281/zenodo.6456585
8. Viviani, J.; Williams, J.T.; Planes, S. (2016). "Two new pygmygobies (Percomorpha: Gobiidae: Trimma) from French Polynesia" (PDF). Journal of the Ocean Science Foundation. 23: 1–11.
9. Suzuki, T.; Sakaue, J.; Senou, H. (2012). "Two New Species of the Gobiid Fish Genus Trimma (Actinopterygii: Perciformes: Gobiidae) from Japan and Palau". Bulletin of the National Museum of Nature and Science. Ser. A (Supplement No. 6): 67–77.
10. Winterbottom, R.; Erdmann, M.V.; Dita Cahyani, N.K. (2014). "Trimma helenae (Pisces; Gobioidei), a new species of gobiid fish from Indonesia" (PDF). Zootaxa. 3760 (3): 420–428. doi:10.11646/zootaxa.3760.3.8. PMID   24870091.
11. Winterbottom, R. (2014). "Trimma irinae, (Pisces; Gobioidei) a new species of gobiid fish from Milne Bay Province, Papua New Guinea" (PDF). Zootaxa. 3802 (2): 209–216. doi:10.11646/zootaxa.3802.2.4. PMID   24871003.
12. Winterbottom, R.; Erdmann, M.V.; Dita Cahyani, N.K. (2014). "Three new species of Trimma (Pisces; Gobioidei) from Indonesia" (PDF). Zootaxa. 3838 (3): 367–384. doi:10.11646/zootaxa.3838.3.7. PMID   25081782.
13. Winterbottom, R.; Pyle, R. L. (2022). "A new species of Trimma (Pisces: Gobiidae) from the deep reefs of Palau, western Pacific Ocean". Zootaxa. 5094 (4): 595–600. doi:10.11646/zootaxa.5094.4.5. PMID   35391433.
14. Hoese, D.F.; Bogorodsky, S.V.; Mal, A.O. (2015). "Description of a new species of Trimma (Perciformes: Gobiidae) from the Red Sea, with a discussion of the generic separation of Trimma and Priolepis, with discussion of sensory papillae terminology" (PDF). Zootaxa. 4027 (4): 538–550. doi:10.11646/zootaxa.4027.4.4. PMID   26624195.
15. Suzuki, T.; Yano, K.; Senou, H. (2015). "Trimma yoshinoi, a new gobiid fish from Japan (Perciformes: Gobiidae)" (PDF). Journal of the Ocean Science Foundation. 14: 66–73.