Williamsoniaceae

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Williamsoniaceae
Temporal range: Triassic–Cretaceous
Williamsonia life restoration.png
Restoration of Wielandiella
Weltrichia diagram.png
Diagram of Weltrichia , a male reproductive structure
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Order: Bennettitales
Family: Williamsoniaceae
(Carruthers) Nathorst, 1913
Genera

Williamsoniaceae is a family within the Bennettitales, an extinct group of seed plants within the Cycadophyta subdivision. Members of this family are believed to have been around two meters tall and with widely serrate leaves along a central stem. Reproductive organs of the Williamsoniaceae have varied widely in the fossil record but almost all have been found to be borne on stalks emerging from a ring of leaves. [1]

Pterophyllum fossil from Shamshak Formation, Middle Jurassic, Iran Pterophyllum fossil from Shamshak Formation, Middle Jurassic, Iran.jpg
Pterophyllum fossil from Shamshak Formation, Middle Jurassic, Iran

Reproduction

This family is different from Cycadeoidaceae by having the presence of cones leaving the major axis and lateral branches associated with a long peduncle covered by bracts. Some of this family reproduce by sporangia and others only produce ovule or pollen sacs. [2]

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<i>Zamites</i> Extinct genus of bennettitalean foliage

Zamites is a genus of sterile foliage known from the Mesozoic of North America, Europe, India and Antarctica through the Eocene of North America. It was erected as a form taxon for leaves that superficially resembled the extant cycad Zamia, however it is now believed to belong to a similar but phylogenetically different group, the cyacadeoids (Bennettitales). The fronds are linear or lanceolate in shape, and pinnately compound, with pinnae with parallel veins and smooth margins, and symmetrical and constricted at the base where they are attached obliquely to the upper surface of the rachis. It has been interpreted as a Bennettitalean plant in the family Williamsoniaceae. It is associated with the ovulate cone Williamsonia and male cone Weltrichia.

<i>Pterophyllum</i> (plant) Extinct genus of seed plants

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Nilssoniopteris is an extinct form genus of leaves belonging to the Bennettitales. Leaves are slender and often entire-margined (smooth-edged), though some species have dissected leaves with numerous small segments extending down to the rachis of the leaf. Nilssoniopteris-like leaves are distinguished by their syndetocheilic stomata, indicating bennettitalean affinities. Similar "taeniopterid" leaves are placed in the genus Nilssonia if their stomata are instead haplocheilic, or Taeniopteris if the cuticle is not preserved. Leaves of Nilssoniopteris vittata from the Middle Jurassic of England are associated with bisexual Williamsoniella reproductive structures.

References

  1. Meyen, Sergei V. (1984). "Basic Features of Gymnosperm Systematics and Phylogeny as Evidenced by the Fossil Record". Botanical Review. 50 (1): 1–111. doi:10.1007/BF02874305. JSTOR   4354028. S2CID   38485693.
  2. Paleontologia Paleovertebrados e Paleobotânica Autor : Ismar de Souza Carvalho. Editora Interciência. Archived 2016-03-04 at the Wayback Machine