Agnostida are an order of extinct arthropods which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician.
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
Agnostus is a genus of agnostid trilobites, belonging to the family Agnostidae, that lived during the late Middle Cambrian – early Upper Cambrian. It is the type genus of the family Agnostidae and is subdivided into two subgenera, Agnostus and Homagnostus.
Acidiscus Rasetti, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida Salter (1864). It lived during the Botomian stage = late Lower Cambrian Stage 4 ; the upper Botomian boundary corresponds to base of the Middle Cambrian, Miaolingian Series and Wuliuan stage.
Acimetopus Rasetti, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi (1943), Order Agnostida Salter (1864). It lived during the Botomian stage. = late Lower Cambrian Stage 4 ; the upper Botomian boundary corresponds to base of the Middle Cambrian, Miaolingian Series and Wuliuan stage.
Analox Rasetti, 1966 is a genus of Eodiscinid trilobites belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida It lived during the Botomian stage. It can easily be distinguished from other trilobites by the two furrows that extend forwards and sidewards from the front of the glabella.
Bolboparia is an extinct genus of eodiscinid agnostid trilobites. It lived during the late Lower Cambrian in what today Canada and the United States.
Cobboldites Kobayashi, 1943, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida It lived during the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the Cambrian Period.
Chelediscus Rushton, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida Salter (1864). The Treatise assigns this genus to the Calodiscidae; Cotton and Fortey (2005) however move it to the Weymouthiidae. Chelediscus lived during the later part of the Botomian stage.
Pagetia is a genus of small trilobite, assigned to the Eodiscinid family Pagetiidae and which had global distribution during the Middle Cambrian. The genus contains 55 currently recognized species, each with limited spatial and temporal ranges.
Yukoniidae S. Zhang, 1980 [nom. transl. et emend. Jell, in Whittington et al., 1997 ex Yukoniinae S. Zhang in W. Zhang, Lu et al., 1980] is a small family of trilobites, belonging to the Eodiscina.
Acmarhachis is a genus of trilobites in the order Agnostida, which lived in what are now Australia, Canada, China (Anhui), Kazakhstan, Russia (Kharaulakh), and the US. It was described by Resser in 1938, and the type species is Acmarhachis typicalis.
Phalagnostus is a genus of small trilobites, in the order Agnostida. It lived during the Middle Cambrian, in what are now Canada, China, the Czech Republic, Denmark, England, France, the Russian Federation, Wales, Sweden, and possibly the United States (Vermont). The headshield is almost entirely effaced and wider than the tailshield. The pygidium is also very effaced, but the ovate pygidial axis is well defined and a border furrow is also present.
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.
Ptychagnostidae is a family of agnostid trilobites from the 5th Stage to the Paibian Age of the Cambrian. The family includes several important index fossils.
Morocconus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It is still debated if it lived at the very end of the Lower Cambrian or at the very beginning of the Middle Cambrian. Morocconus notabilis is the only known species in this genus.
The Peronopsidae comprise the earliest family of the Agnostina suborder. Species of this family occurred on all paleocontinents. The earliest representatives of this family first occur just before the start of the Middle Cambrian, and the last disappeared just after the start of the Upper Cambrian.
The Condylopygidae Raymond (2013) are a family of small trilobites that lived during the Middle Cambrian, and found in Canada, the Czech Republic, Germany, France, Spain, England, Wales, Sweden, and the Russian Federation (Siberia). They uniquely differ from all other Agnostina in having the frontal glabellar lobe wider than the rear lobe. The Condylopygidae are the only family assigned to the Condylopygoidea superfamily.
Cyclopyge is a genus of small to average size trilobites that lived during the Ordovician. Like all members of the family Cyclopigidae, it has very large convex eyes, that cover most of the free cheeks, and in some species touch each other. The eyes almost touch the large glabella. The occipital ring has merged with the rest of the glabella. The glabella does not extend into a frontal thorn. The cephalon lacks genal spines. The 6 thorax segments have short pleurae. The pygidium is rather large, and often rather effaced. These are features that also occur in other Cyclopygidae, and are indications of a pelagic lifestyle.
Librostoma is a subclass of trilobites defined by having a natant hypostome, which is a hypostome that is free from the anterior doublure and aligned with the anterior of the glabella, this is unlike a conterminant hypostome, which is attached to the exoskeleton.
