Agraulos Temporal range: Middle Cambrian ~ | |
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Agraulos ceticephalus Menevia Formation, Wales. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | † Trilobita |
Order: | † Ptychopariida |
Family: | † Solenopleuridae |
Subfamily: | † Agraulinae |
Genus: | † Agraulos Hawle & Corda, 1847 |
Species | |
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Synonyms | |
Agraulos synonymy
A. ceticephalus synonymy
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Agraulos is a genus of Solenopleuridae [1] trilobites that lived during the Middle Cambrian in North America and Europe, particularly the Czech Republic. The genus was named by Hawle & Corda in 1847. [2]
Agraulos is derived from the Greek Ἄγραυλος, "country woman", wife of Kekrops. [3]
Type species (designated by Miller 1889). [4] Arion ceticephalus Barrande, 1846 [5] from the Cambrian Eccaparadoxides pusillus Zone in the Skryje Member of the Buchava Formation, within the Skryje–Tyrovice Basin, Bohemia.
Agraulinae with cephala generally domed; glabella isosceles-trapezoidal, i.e. with truncate front and base angles of the forward-converging lateral margins/flanks more than 15°; occipital ring mesially swollen backwards, with or without a medial node or spine; preglabellar field relatively long (sag.); posterolateral projection of fixigena narrow (tr.); librigenal spines short to long, with some deflected outwards. Thorax of up to 16 segments with first anterior axial rings marked by terrace lines immediately succeeded in some species by rings bearing incipient median nodes or incipient/prominent spines; thoracic segments finely punctate or granulate. Pygidium, small and transverse (Fletcher, 2017, pp, 9,10).
Arionellus quadrangularis Whitfield (1884), originally collected from the mid-Cambrian Braintree Formation at Old Hayward Quarry, Quincy, Massachusetts, US, had previously been assigned to Agraulos by Walcott and others. [18] McMenamin (2002), however, erected Skehanos to accommodate the species as Skehanos quadrangularis . [19]
Scandinavian species attributed by Westergård (1953) [20] to Agraulos [i.e. A. difformis, A. aculeatus, A. acuminatus (all Angelin, 1851), and A. anceps] were transferred by Ahlberg & Bergström (1978) [21] to Proampyx [22] although several Lower Cambrian forms they also assigned to Proampyx would later be allocated to various other genera.
Paradoxides is a genus of large to very large trilobite found throughout the world during the Middle Cambrian period. One record-breaking specimen of Paradoxides davidis, described by John William Salter in 1863, is 37 cm (15 in). The cephalon was semicircular with free cheeks ending in long, narrow, recurved spines. Eyes were crescent shaped providing an almost 360° view, but only in the horizontal plane. Its elongate thorax was composed of 19–21 segments and adorned with longish, recurved pleural spines. Its pygidium was comparatively small. Paradoxides is a characteristic Middle-Cambrian trilobite of the 'Atlantic' (Avalonian) fauna. Avalonian rocks were deposited near a small continent called Avalonia in the Paleozoic Iapetus Ocean. Avalonian beds are now in a narrow strip along the East Coast of North America, and in Europe.
Agnostus is a genus of agnostid trilobites, belonging to the family Agnostidae, that lived during the late Middle Cambrian – early Upper Cambrian. It is the type genus of the family Agnostidae and is subdivided into two subgenera, Agnostus and Homagnostus.
The Paradoxididae are a family of trilobites, a group of extinct marine arthropods. They occurred during the late Lower Cambrian (Toyonian) and disappeared at the end of the Middle Cambrian. Representatives of this family have been found in the paleocontinents of Avalonia, Baltica, and Gondwana, now Canada, USA, England, Wales, Morocco, Spain, Czech Republic, Poland, Russia, Mongolia, and Turkey. Species in this family can typically grow large to very large, are relatively flat, have an inverted egg-shaped outline, opisthoparian sutures, a glabella that in early genera has parallel sides and expands forward in later representatives, and approaches or reaches the frontal border. All species have an almost semicircular headshield with long backward-directed genal spines. The articulate middle part of the body consists of 15 to 21 segments ending in sickle-shaped spines that to the back curve increasingly further backwards. The tailshield is small.
Acontheus is a genus of trilobites belonging to the Family Corynexochidae, Order Corynexochida, and is geographically widespread having been recorded from middle Cambrian strata in Sweden, Newfoundland, Germany, Siberia, Antarctica, Queensland, China and Wales.
Acidiscus Rasetti, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida Salter (1864). It lived during the Botomian stage = late Lower Cambrian Stage 4 ; the upper Botomian boundary corresponds to base of the Middle Cambrian, Miaolingian Series and Wuliuan stage.
Chelediscus Rushton, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida Salter (1864). The Treatise assigns this genus to the Calodiscidae; Cotton and Fortey (2005) however move it to the Weymouthiidae. Chelediscus lived during the later part of the Botomian stage.
Wanneria is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the later part of the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the Cambrian Period. Wanneria walcottana is the only known species in this genus.
Peronopsis is a genus of trilobite restricted to the Middle Cambrian. Its remains have been found in Asia, Australia, Europe, and North America.
The Wheeler Shale is a Cambrian (c. 507 Ma) fossil locality world-famous for prolific agnostid and Elrathia kingii trilobite remains and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna and preservation style normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten.
Pagetia Walcott, 1916. is a small genus of trilobite, assigned to the Eodiscinid family Pagetiidae and which had global distribution during the Middle Cambrian. The genus contains 55 currently recognized species, each with limited spatial and temporal ranges.
Ellipsocephalus Zenker, 1833, is a genus of blind Cambrian trilobite, comprising benthic species inhabiting deep, poorly lit or aphotic habitats. E. hoffi is a common trilobite mainly from central Europe.
Phalagnostus is a genus of small trilobites, in the order Agnostida. It lived during the Middle Cambrian, in what are now Canada, China, the Czech Republic, Denmark, England, France, the Russian Federation, Wales, Sweden, and possibly the United States (Vermont). The headshield is almost entirely effaced and wider than the tailshield. The pygidium is also very effaced, but the ovate pygidial axis is well defined and a border furrow is also present.
Lejopyge laevigata is a species of agnostid trilobite belonging to the genus Lejopyge. It existed during the Guzhangian to the Paibian Age of the Cambrian. It has a cosmopolitan distribution and is an important index fossil in biostratigraphy.
Pleuroctenium Hawle & Corda (1847) is an agnostid trilobite belonging to the family Condylopygidae Raymond (1913). The genus occurs in Middle Cambrian (Drumian) strata of Canada, the Czech Republic, England and Wales, France, and Sweden.
Diplorrhina Hawle and Corda (1847) is a genus of trilobite belonging to Order Agnostida. It lived during the early Middle Cambrian in what are now the Czech Republic and the North Siberian plateau. as in members of the family Peronopsidae it lacks a preglabellar furrow. Both cephalon and pygidium lack spines. It is difficult to distinguish Diplorrhina from many other peronopsids.
Condylopyge Hawle and Corda (1847) is a genus of agnostid trilobite that lived during the late Lower and early Middle Cambrian, in what are today Canada, the Czech Republic, England and Wales, France, Germany, Italy, Morocco, the Russian Federation, Spain, Turkey and Sweden. It can easily be distinguished from all other Agnostida because the frontal glabellar lobe is notably wider than the rear lobe. It belongs to the same family as Pleuroctenium but the frontal glabellar lobe does not fold around the rear lobe, as it does in that genus. Condylopyge is long ranging, possibly spanning the early Cambrian Terreneuvian Series in Nuneaton, central England into at least Drumian strata at various locations elsewhere.
Tannudiscus Pokrovskaya (1959) is a genus of Eodiscinid trilobites belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida. It lived during the late Lower Cambrian, with remains found in Canada (Newfoundland), China (Gansu), The United Kingdom (England), and the Russian Federation.
Plutonides is a genus of Middle Cambrian trilobite in the family Paradoxididae with species Plutonides sedgwickii and possibly Plutonides? illingi. Several other species and subspecies were at times placed in Plutonides but have subsequently been moved to other genera.
Serrodiscus Richter and Richter 1941. is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida. It lived during the late Lower Cambrian, with remains found in Canada, China (Gansu), The United Kingdom (England), Germany (Silesia), Poland, the Russian Federation, and the United States. It is named for the spines on the ventral side of the pygidium, which give it a serrated impression.
Bailiaspis Resser, 1936, is a Middle Cambrian (Miaolingian) trilobite genus belonging to the Family Conocoryphidae Angelin, 1854. Within the Acado-Baltic region, the genus ranges from Wuliuan into Guzhangian age strata.