Anatoma

Anatoma
Two views of a shell of Anatoma umbilicata
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Vetigastropoda
Order:	Lepetellida
Superfamily:	Scissurelloidea
Family:	Anatomidae
Genus:	Anatoma Woodward, 1859
Type species
A. crispata Fleming, 1828  [1]
Synonyms [2]
AnatomusH. & A. Adams HainellaBandel, 1998 SchizotrochusMonterosato, 1877 Scissurella (Anatoma)Woodward, 1859 (original rank) Scissurella (Schizotrochus)Monterosato, 1817 junior subjective synonym ThieleellaBandel, 1998

Anatoma is a genus of minute marine gastropod molluscs or micromolluscs in the family Anatomidae, found in Europe, Australia and New Zealand. ^[2]

They were formerly placed as a subfamily in the family Scissurellidae. They were raised to family level by Geiger and Jansen in 2004. ^[3]

Description

Anatoma snails have conical or top-shaped (turbiniform) shells. The shell often features a slit or hole in the margin of the outer lip and possesses a deep, perspective umbilicus. This genus is further characterized a rhipidoglossate radula, and a lack of nacre. Their shells may be sculptured with longitudinal lamellae and transverse lines, along with ornamentation like concentric elevated lirae. ^[4]

Species

The genus Anatoma includes the following species: ^[2]

• Anatoma aedonia (Watson, 1886)
• Anatoma africana (Barnard, 1963)
• Anatoma agulhasensis (Thiele, 1925)
• Anatoma alta (Watson, 1886)
• Anatoma alternatisculpta Geiger & McLean, 2010
• Anatoma amoena (Thiele, 1912)
• Anatoma amydra Geiger & B.A. Marshall, 2012
• Anatoma argentinae (Zelaya & Geiger, 2007)
• Anatoma aspera (Philippi, 1844)
• Anatoma atlantica (Bandel, 1998)
• Anatoma aupouria (Powell, 1937)
• Anatoma australis (Hedley, 1903)
• Anatoma austrolissa Geiger & Sasaki, 2008
• Anatoma balgimae Utrilla & Gofas, 2024
• Anatoma bathypacifica (Geiger & McLean, 2010)
• Anatoma biconica Geiger, 2012
• Anatoma bisculpta L. Hoffman, Gofas & Freiwald, 2021
• Anatoma boucheti Geiger & Sasaki, 2008
• Anatoma breveprima Geiger, 2012
• Anatoma brychia Pimenta & Geiger, 2015
• Anatoma campense Pimenta & Geiger, 2015
• Anatoma concinna A. Adams, 1862
• Anatoma conica (d'Orbigny, 1841)
• Anatoma copiosa Pimenta & Geiger, 2015
• Anatoma corralae Serge GOFAS, Ángel A. LUQUE, Joan Daniel OLIVER, José TEMPLADO & Alberto SERRA (2021), 2021 ^[5]
• Anatoma crispata (Fleming, 1828)
• Anatoma declivis L. Hoffman, Kniesz, Martínez Arbizu & Kihara, 2022
• Anatoma discapex L. Hoffman, Kniesz, Martínez Arbizu & Kihara, 2022
• Anatoma disciformis (Golikov & Sirenko, 1980)
• Anatoma emilioi Geiger, 2011 ^[6]
• Anatoma equatoria (Hedley, 1899)
• Anatoma espiritosantense Pimenta & Geiger, 2015
• Anatoma euglypta (Pelseneer, 1903)
• Anatoma eximia (Seguenza, 1877)
• Anatoma finlayi (Powell, 1937)
• Anatoma flemingi (B.A. Marshall, 2002)
• Anatoma flexidentata Geiger & Sasaki, 2008
• Anatoma fujikurai Sasaki, Geiger & Okutani 2010
• Anatoma georgii Geiger, 2017
• † Anatoma gephrya Maxwell, 1992
• Anatoma globulus Geiger, 2012
• Anatoma golikovi Nekhaev & Krol, 2020
• Anatoma gunteri (Cotton & Godfrey, 1933)
• † Anatoma hedegaardi (Bandel, 1998)
• Anatoma hyposculpta Geiger, 2012
• Anatoma indonesica Bandel, 1998
• Anatoma janetae Geiger, 2006
• Anatoma janusa Geiger, 2012
• Anatoma japonica (A. Adams, 1862)
• Anatoma keenae (McLean, 1970)
• Anatoma kelseyi (Dall, 1905)
• Anatoma kopua Geiger & B.A. Marshall, 2012
• Anatoma laevapex L. Hoffman, Kniesz, Martínez Arbizu & Kihara, 2022
• Anatoma lamellata (A. Adams, 1862)
• Anatoma lyra (S. S. Berry, 1947)
• Anatoma maxima (Schepman, 1908)
• Anatoma megascutula Geiger & B.A. Marshall, 2012
• Anatoma micalii Geiger, 2012
• † Anatoma miocenica (Laws, 1939)
• Anatoma munieri (P. Fischer, 1862)
• Anatoma occidentalis L. Hoffman & Freiwald, 2022
• Anatoma orbiculata Geiger, 2012
• Anatoma pagoda L. Hoffman, Gofas & Freiwald, 2021
• Anatoma parageia Geiger & Sasaki, 2009
• Anatoma paucisculpta L. Hoffman, Kniesz, Martínez Arbizu & Kihara, 2022
• Anatoma peruviana (Geiger & McLean, 2010)
• † Anatoma philippiana (O. Semper, 1865)
• Anatoma philippinica (Bandel, 1998)
• Anatoma planapex Geiger, 2012
• Anatoma plicatazona Geiger & McLean, 2010
• Anatoma porcellana Geiger, 2012
• Anatoma proxima (Dall, 1927)
• Anatoma pseudoequatoria (Kay, 1979)
• Anatoma quadraxialis Geiger, 2012
• Anatoma rainesi Geiger, 2003
• Anatoma rapaensis Geiger, 2008
• † Anatoma redoniana Landau, Van Dingenen & Ceulemans, 2017
• Anatoma regia (Mestayer, 1916)
• Anatoma rhynchodentata Geiger, 2012
• Anatoma richardi (Dautzenberg & H. Fischer, 1896)
• Anatoma rolani Geiger & Fernández-Garcés, 2010
• † Anatoma rosenkrantzi Schnetler, Lozouet & Pacaud, 2001
• Anatoma sagamiana (Okutani, 1964)
• Anatoma schanderi Høisæter & Geiger, 2011
• Anatoma shiraseae Numanami & Okutani, 1990
• Anatoma sinuosa Geiger, 2012
• Anatoma soyoae (Habe, 1951)
• Anatoma symmetrica L. Hoffman, Gofas & Freiwald, 2021
• Anatoma tabulata (Barnard, 1964)
• Anatoma tangaroa Geiger & B.A. Marshall, 2012
• Anatoma tenuis (Jeffreys, 1877)
• Anatoma tenuisculpta (Seguenza, 1877)
• Anatoma tobeyoides Geiger & Jansen, 2004
• † Anatoma tongaensis (Ladd, 1970)
• Anatoma umbilicata (Jeffreys, 1883)
• Anatoma weddelliana (Zelaya & Geiger, 2007)
• Anatoma yaroni Herbert, 1986
• Anatoma zancliformis Geiger, 2012

Synonyms include: ^[2]

• Anatoma aequatorinaHedley, 1899: synonym of Anatoma equatoria (Hedley, 1899)
• Anatoma aetheria(Melvill & Standen, 1903): synonym of Anatoma japonica(A. Adams, 1862)
• Anatoma americanaBandel, 1998: synonym of Anatoma proxima (Dall, 1927)
• Anatoma baxteriMcLean, 1984: synonym of Thieleella baxteri (McLean, 1984)
• Anatoma cebuanaBandel, 1998: synonym of Scissurella cebuana (Bandel, 1998)
• Anatoma chiricova(Dall, 1919): synonym of Anatoma kelseyi (Dall, 1905)
• † Anatoma costamagnaensis Bandel, 1998: synonym of Anatoma aspera (Philippi, 1844)
• Anatoma dohrnianaDunker, 1861: synonym of Scissurella dohrniana (Dunker, 1861)
• Anatoma epicharis(McLean, 1970): synonym of Anatoma keenae (McLean, 1970)
• Anatoma exquisita(Schepman, 1908): synonym of Anatoma japonica (A. Adams, 1862)
• Anatoma funiculataGeiger & Jansen, 2004: synonym of Anatoma pseudoequatoria (Kay, 1979)
• Anatoma herbertiGeiger & Sasaki, 2008: synonym of Anatoma finlayi (Powell, 1937)
• Anatoma jacksoni(Melvill, 1904): synonym of Anatoma munieri (P. Fischer, 1862)
• Anatoma jansenaeGeiger, 2006: synonym of Anatoma japonica (A. Adams, 1862)
• Anatoma josephinae(Odhner, 1960): synonym of Anatoma tenuis (Jeffreys, 1877)
• Anatoma mantelli(Woodward, 1859): synonym of Scissurella mantelli Woodward, 1859
• Anatoma mirificaA. Adams, 1862: synonym of Scissurella mirifica (A. Adams, 1862)
• Anatoma obtusata(Golikov & Gulbin, 1978): synonym of Anatoma concinna (A. Adams, 1862)
• Anatoma paucispiralia(Bandel, 1998): synonym of Anatoma pseudoequatoria (Kay, 1979)
• Anatoma pulchella(Bandel, 1998): synonym of Anatoma pseudoequatoria (Kay, 1979)
• Anatoma schioetteiHøisæter & Geiger, 2011: synonym of Anatoma crispata (J. Fleming, 1828) (a junior synonym)
• Anatoma stamineaA. Adams, 1862: synonym of Scissurella staminea (A. Adams, 1862)
• Anatoma turbinataA. Adams, 1862: synonym of Anatoma munieri (P. Fischer, 1862)

Nomen dubium

• Anatoma bertheloti(d'Orbigny, 1840)

References

1. Daniel L. Geiger; Takenori Sasaki (2008). "Four new species of Anatomidae (Mollusca: Vetigastropoda) from the Indian Ocean (Reunion, Mayotte) and Australia, with notes on a novel radular type for the family" (PDF). Zoosymposia . 1: 247–264. doi: 10.11646/zoosymposia.1.1.14 .
2. Bouchet, P.; Gofas, S. (2011). Anatoma Woodward, 1859. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138464 on 2012-01-14,
3. Geiger D.L. & Jansen P. (2004). New species of Australian Scissurellidae (Mollusca: Gastropoda: Vetigastropoda) with remarks on Australian and Indo-Malayan species. Zootaxa 714:1-72.
4. Geiger, D. L., & Marshall, B. A. (2012). New species of Anatomidae (Mollusca: Gastropoda: Vetigastropoda) from New Zealand, and new data on fourayas species of Anatomidae and one species of Scissurellidae. Zootaxa, 3484, 1-22.
5. Serge GOFAS, Ángel A. LUQUE, Joan Daniel OLIVER, José TEMPLADO & Alberto SERRA (2021) - The Mollusca of Galicia Bank (NE Atlantic Ocean); European Journal of Taxonomy 785: 1–114
6. Daniel L. Geiger (2011). Anatoma emilioi, a new species of Anatomidae McLean, 1989 from the Hawaiian Islands. - 534-535. In: Mike Severns. Shells of the Hawaiian Islands - The Sea Shells: 564 pp., ConchBooks, Hackenheim.

• Woodward S. P. (1859). On a new species of mollusk of the genus Scissurella D'Orbigny. Proceedings of the Zoological Society of London 27 (392-405): 202-204
• Høisæter T. & Geiger D.L. (2011) Species of Anatoma (Gastropoda: Anatomidae) in Norwegian and adjacent waters, with the description of two new species. The Nautilus 125(3): 89-112.

Further reading

• Geiger, D. L. (2003). "Phylogenetic assessment of characters proposed for the generic classification of Recent Scissurellidae (Gastropoda: Vetigastropoda) with a description of one new genus and six new species from Easter Island and Australia". Molluscan Research . 23 (1): 21–83. doi: 10.1071/MR02017 .
• Geiger, D. L. & McLean, J. H. (2010). "New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda)" (PDF excerpt). Zootaxa . 2356: 1–35, 24 plates. doi:10.11646/zootaxa.2356.1.1.
• Powell, A. W. B. (1979). New Zealand Mollusca. Auckland, New Zealand: William Collins Publishers Ltd. ISBN   0-00-216906-1.
• Geiger D.L. (2012) Monograph of the little slit shells. Volume 1. Introduction, Scissurellidae. pp. 1-728. Volume 2. Anatomidae, Larocheidae, Depressizonidae, Sutilizonidae, Temnocinclidae. pp. 729-1291. Santa Barbara Museum of Natural History Monographs Number 7.