The Carboniferous is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period 358.9 million years ago (mya), to the beginning of the Permian Period, 298.9 mya. The name Carboniferous means "coal-bearing", from the Latin carbō ("coal") and ferō, and refers to the many coal beds formed globally during that time.
Dictyoptera is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea and the order Mantodea (mantises). All modern Dictyoptera have short ovipositors and typically lay oothecae. The oldest fossils of Dictyoptera from the Late Carboniferous, referred to as "roachoids" have long ovipositors and did not lay oothecae. The oldest modern oothecae-laying dictyopterans date to the Late Triassic.
Meganeura is a genus of extinct insects from the Late Carboniferous. They resembled and are related to the present-day dragonflies and damselflies, and were predatory, with their diet mainly consisting of other insects. The genus belongs to the Meganeuridae, a family including other similarly giant dragonfly-like insects ranging from the Late Carboniferous to Middle Permian. With a wingspan ranging from 65 cm (25.6 in) to over 70 cm (28 in), M. monyi is one of the largest-known flying insect species.
Holometabolism, also called complete metamorphosis, is a form of insect development which includes four life stages: egg, larva, pupa, and imago. Holometabolism is a synapomorphic trait of all insects in the superorder Endopterygota. Immature stages of holometabolous insects are very different from the mature stage. In some species the holometabolous life cycle prevents larvae from competing with adults because they inhabit different ecological niches. The morphology and behavior of each stage are adapted for different activities. For example, larval traits maximize feeding, growth, and development, while adult traits enable dispersal, mating, and egg laying. Some species of holometabolous insects protect and feed their offspring. Other insect developmental strategies include ametabolism and hemimetabolism.
The American cockroach is the largest species of common cockroach, and often considered a pest. In certain regions of the U.S. it is colloquially known as the waterbug, though it is not a true waterbug since it is not aquatic. It is also known as the ship cockroach, kakerlac, and Bombay canary. It is often misidentified as a palmetto bug.
The Madagascar hissing cockroach, also known as the hissing cockroach or simply hisser, is one of the largest species of cockroach, reaching 5 to 7.5 centimetres at maturity. They are native to the island of Madagascar, which is off the African mainland, where they are commonly found in rotting logs. It is one of some 20 known species of large hissing roaches from Madagascar, many of which are kept as pets, and often confused with one another by pet dealers; in particular, G. portentosa is commonly confused with G. oblongonota and G. picea.
Blattodea is an order of insects that contains cockroaches and termites. Formerly, termites were considered a separate order, Isoptera, but genetic and molecular evidence suggests they evolved from within the cockroach lineage, cladistically making them cockroaches as well. The Blattodea and the mantis are now all considered part of the superorder Dictyoptera. Blattodea includes approximately 4,400 species of cockroach in almost 500 genera, and about 3,000 species of termite in around 300 genera.
Cockroaches are insects belonging to the order Blattaria. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known as pests.
The Palaeodictyoptera are an extinct order of medium-sized to very large, primitive Palaeozoic paleopterous insects. They are informative about the evolution of wings in insects.
Meganisoptera is an extinct order of very large to gigantic winged insects, informally known as griffinflies or (incorrectly) as giant dragonflies. The order was formerly named Protodonata, the "proto-Odonata", for their similar appearance and supposed relation to modern Odonata. They range in Palaeozoic times. Though most were only slightly larger than modern dragonflies, the order includes the largest known insect species, such as the late Carboniferous Meganeura monyi and the even larger early Permian Meganeuropsis permiana, with wingspans of up to 71 centimetres (28 in).
The most recent understanding of the evolution of insects is based on studies of the following branches of science: molecular biology, insect morphology, paleontology, insect taxonomy, evolution, embryology, bioinformatics and scientific computing. It is estimated that the class of insects originated on Earth about 480 million years ago, in the Ordovician, at about the same time terrestrial plants appeared. Insects are thought to have evolved from a group of crustaceans. The first insects were landbound, but about 400 million years ago in the Devonian period one lineage of insects evolved flight, the first animals to do so. The oldest insect fossil has been proposed to be Rhyniognatha hirsti, estimated to be 400 million years old, but the insect identity of the fossil has been contested. Global climate conditions changed several times during the history of Earth, and along with it the diversity of insects. The Pterygotes underwent a major radiation in the Carboniferous while the Endopterygota underwent another major radiation in the Permian.
Apthoroblattina is an extinct genus of primitive cockroaches from the Carboniferous period. Fossils of the genus have been found in England, Wales, the United States, and Russia. The paratype specimen for the species, A. johnsoni, is recorded to have a total length of 43 mm and a width of 38 mm, while the type specimens of A. sulcata is noted to have been up to 45 mm in length and 25 mm in width if complete.
Writhlington SSSI is a 0.5 hectare geological Site of Special Scientific Interest near the town of Radstock, Bath and North East Somerset, notified in 1992.
"Roachoids", also known as "Roachids", "Blattoids" or Eoblattodea, are members of the stem group of Dictyoptera. They generally resemble cockroaches, but most members, unlike modern dictyopterans, have generally long external ovipositors, and are thought not to have laid ootheca like modern dictyopterans.
Idelinellidae is a family of fossil cockroach-like insects. They closely resemble members of the family Stenoneuridae, and consist of the following species:
Paleontology in Rhode Island refers to paleontological research occurring within or conducted by people from the U.S. state of Rhode Island. Rhode Island has a relatively sparse fossil record. Among its more common fossils are plant remains that are frequently associated with the state's coal beds. During the early Paleozoic, Rhode Island was at least partially submerged under a sea inhabited by trilobites. During the Carboniferous period the state became a swampy environment where lush vegetation included trees more than 50 feet high. Local animal life included arachnids and insects like cockroaches. Rift basins formed locally during the Permian. The ensuing Triassic and Jurassic periods are absent from the state's rock record. Little is known about the state's Cretaceous history. The Paleogene and Neogene periods are also missing from Rhode Island's rock record. During the Pleistocene the state was subjected to glacial activity. Notable local fossil finds have included previously unknown kinds of insect and abundant ancient amphibian trackways.
Blattoidea is a superfamily of cockroaches and termites in the order Blattodea. There are about 17 families and more than 4,100 described species in Blattoidea.
Mazothairos is an extinct genus of very large insect from the Carboniferous period. It was a member of the order Palaeodictyoptera. Although it is only known from very fragmentary remains from a single fossil, it is estimated to have had a wingspan of about 56 centimeters (22 in), making it one of the largest-known insects, only being rivaled in size by the largest members of the order Meganisoptera, such as Meganeura and Meganeuropsis.
