Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.
Paradoxides is a genus of large to very large trilobite found throughout the world during the Middle Cambrian period. One record-breaking specimen of Paradoxides davidis, described by John William Salter in 1863, is 37 cm (15 in). The cephalon was semicircular with free cheeks ending in long, narrow, recurved spines. Eyes were crescent shaped providing an almost 360° view, but only in the horizontal plane. Its elongate thorax was composed of 19–21 segments and adorned with longish, recurved pleural spines. Its pygidium was comparatively small. Paradoxides is a characteristic Middle-Cambrian trilobite of the 'Atlantic' (Avalonian) fauna. Avalonian rocks were deposited near a small continent called Avalonia in the Paleozoic Iapetus Ocean. Avalonian beds are now in a narrow strip along the East Coast of North America, and in Europe.
Olenoides was a trilobite from the Cambrian period. Its fossils are found well-preserved in the Burgess Shale in Canada. It grew up to 10 cm long.
Naraoia is a genus of small to average size marine arthropods within the family Naraoiidae, that lived from the early Cambrian to the late Silurian period. The species are characterized by a large alimentary system and sideways oriented antennas.
Dikelocephalus is a genus of very large trilobites of up to 50 cm (20 in) long, that lived during the last 3 million years of the Cambrian (Sunwaptan). Their fossils are commonly found as disarticulated sclerites, in the upper Mississippi Valley and in Canada (Alberta). The exoskeleton is rounded anteriorly, with the thorax and sides of the tailshield slightly tapering to about 2⁄3× of the width across the base of the spines at the back of the headshield. At the side corners of the pygidium there may be triangular or hooked spines, pointing backwards, while between the spines the posterior margin is at a 30-75° angle with the lateral margin, gently convex or nearly straight. If pygidial spines are lacking, the margin is gradually rounded. The thorax has 12 segments.
Yunnanocephalus is a genus of ptychopariid trilobite. It lived during the late Atdabanian and Botomian stages, in what are currently Antarctica, Australia and China. It was a "moderately common" member of the Chengjiang Fauna. Yunnanocephalus is the only genus currently assigned to the Yunnanocephalidae family.
Meteoraspis is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae. The various species lived from 501 to 497 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Meteoraspis are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as M. nevensis from Victoria Land, Antarctica.
The Wheeler Shale is a Cambrian (c. 507 Ma) fossil locality world-famous for prolific agnostid and Elrathia kingii trilobite remains and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna and preservation style normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten.
The Marjum Formation is a Cambrian geological formation that overlies the Wheeler Shale in the House Range, Utah. It is named after its type locality, Marjum Pass, and was defined in 1908. The formation is known for its occasional preservation of soft-bodied tissue, and is slightly younger than the Burgess Shale, falling in the Ptychagnostus praecurrens trilobite zone.
Alokistocaridae is a family of ptychopariid trilobites that lived from the Botomian epoch of the Early Cambrian until the Late Cambrian. Alokistocarids were particle feeders and left small furrows which are occasionally preserved. Their remains are found worldwide. Elrathia kingii, one of the most collected trilobites in the world, is a typical alokistocarid.
Bathyuriscus is an extinct genus of Cambrian trilobite. It was a nektobenthic predatory carnivore. The genus Bathyuriscus is endemic to the shallow seas that surrounded Laurentia. Its major characteristics are a large forward-reaching glabella, pointed pleurae or pleurae with very short spines, and a medium pygidium with well-impressed furrows. Complete specimens have never reached the size of 7 cm predicted by the largest pygidium found. Bathyuriscus is often found with the free cheeks shed, indicating a moulted exoskeleton. An average specimen will in addition have a furrowed glabella, crescent-shaped eyes, be semi-circular in overall body shape, have 7 to 9 thoracic segments, and a length of about 1.5 inches.
Lotagnostus is a genus of very small trilobites in the order Agnostida, which lived on the outer continental shelves worldwide, during the late Upper Cambrian. It was described by Whitehouse in 1936, and the type species is Lotagnostus trisectus, which was originally described as a species of Agnostus by Salter in 1864.
Ptychoparia is a genus of ptychopariid trilobites, and is the type genus of the family Ptychopariidae, and the order Ptychopariida.
Thoracocare is a minute to very small trilobite, that lived during part of the Middle Cambrian in what are today the states of Idaho, Nevada and Utah. It is the only trilobite known with just two thorax segments outside most members of the Agnostida order. It can be distinguished from Agnostida by the very wide subquadrate glabella, parallel-side or widening forward in the largest specimen, with the full front side touching the border. Two species are known, one, T. idahoensis, only from pygidia.
Cedaria is an extinct genus of trilobites from the late Cambrian.
Kendallina is a genus of parabolinoidid trilobite with an inverted egg-shaped outline, a wide headshield, small eyes, small deflected spines, 12 thorax segments and a small, short tailshield. It lived during the Late Cambrian in what are today Canada and the United States.
Orygmaspis is a genus of asaphid trilobite with an inverted egg-shaped outline, a wide headshield, small eyes, long genal spines, 12 spined thorax segments and a small, short tailshield, with four pairs of spines. It lived during the Late Cambrian in what are today Canada and the United States.
Genevievella is a genus of trilobites with a short inverted egg-shaped outline, a wide headshield, small eyes, and long genal spines. The backrim of the headshield is inflated and overhangs the first of the 9 thorax segments. The 8th thorax segment from the front bears a backward directed spine that reaches beyond the back end of the exoskeleton. It has an almost oval tailshield with 5 pairs of pleural furrows. It lived during the Upper Cambrian in what are today Canada and the United States.
Tricrepicephalus is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae with species of average size. Its species lived from 501 to 497 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Tricrepicephalus are widespread in Late Cambrian deposits in North America, but is also known from one location in South America. Tricrepicephalus has an inverted egg-shaped exoskeleton, with three characteristic pits in the fold that parallels the margin of the headshield just in front of the central raised area. The articulating middle part of the body has 12 segments and the tailshield carries two long, tubular, curved pygidial spines that are reminiscent of earwig's pincers that rise backwards from the plain of the body at approximately 30°.
