Camponotus herculeanus

Last updated

Camponotus herculeanus
Camponotus herculeanus casent0173157 profile 1.jpg
Camponotus herculeanus worker
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Formicinae
Genus: Camponotus
Species:
C. herculeanus
Binomial name
Camponotus herculeanus

Camponotus herculeanus (or Hercules ant) [1] is a species of ant in the genus Camponotus , the carpenter ants, [2] occurring in Northern Eurasia, from Norway to Eastern Siberia, and North America. First described as Formica herculeana by Linnaeus in 1758, [3] the species was moved to Camponotus by Mayr in 1861. [4]

Contents

Head of a queen dealate Camponotus ligniperda.jpg
Head of a queen dealate

Description

The colony of Camponotus herculeanus consists of one or several wingless females (queens), some fertile males, and three castes of sterile workers, known as majors, intermediates, and minors, in decreasing order of size. The queens are large, about 15 mm (0.6 in) in length, and are blackish in colour. The males are a similar colour but about half the size of the queens. The workers usually have blackish heads and gasters, and dark reddish-brown mesosomas, petioles and legs. In majors, the scapes (the long segments of the antenna, before the elbow) are shorter than the length of the head; in intermediates they are about the same length, and in minors, they extend well beyond the back of the head. The head and the dorsal surfaces of the mesosoma and gaster of the largest majors are bristly. [5]

Distribution and habitat

Camponotus herculeanus has a widespread distribution in the Northern Hemisphere, being present in most of Europe, Central and Northern Asia, Canada and the United States. It is common in mountainous regions and is the dominant ant species in mountainous and northerly parts of North America. It occupies a range of habitats including various types of conifer and hardwood forests, clearings, oak scrubland, disturbed areas, pastures and seashore grassland. [5]

Ecology

Nests of Camponotus herculeanus are built in timber, living or rotting trees, stumps, fallen logs and occasionally the structural timbers of buildings. The ants use their strong jaws to excavate galleries and chambers under the bark or in the wood, with a preference for damp wood or timber with fungal decay. In standing trees, their tunnels sometimes extend for 10 m (30 ft) above the ground. [1] Satellite colonies, linked to the original nest by underground tunnels, may develop nearby, often in warmer, drier locations. These house older larvae, pupae, winged reproductives and workers, with the eggs and younger larvae remaining in the main nest. [6]

A colony of Camponotus herculeanus contains several wingless females, which may be unrelated. Winged reproductives are produced in late summer and overwinter in the colony, emerging to fly in swarms on warm spring days. The workers become active in spring and forage in the vicinity of the nest. [5] They tend aphids, and the larvae of the silvery blue butterfly (Glaucopsyche lygdamus), which often feeds on the lupine Lupinus bakeri . [5] The diet consists of the honeydew produced by sap-sucking insects and the ants also consume any insect larvae that they encounter. [7] The ant cricket Myrmecophilus pergandei sometimes lives in the colony, where it is tolerated by the ants. [5]

Related Research Articles

<span class="mw-page-title-main">Ant</span> Family of insects

Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22,000 species have been classified. They are easily identified by their geniculate (elbowed) antennae and the distinctive node-like structure that forms their slender waists.

<span class="mw-page-title-main">Weaver ant</span> Genus of ants

Weaver ants or green ants are eusocial insects of the Hymenoptera family Formicidae belonging to the tribe Oecophyllini. Weaver ants live in trees and are known for their unique nest building behaviour where workers construct nests by weaving together leaves using larval silk. Colonies can be extremely large consisting of more than a hundred nests spanning numerous trees and containing more than half a million workers. Like many other ant species, weaver ants prey on small insects and supplement their diet with carbohydrate-rich honeydew excreted by scale insects (Hemiptera). Weaver ant workers exhibit a clear bimodal size distribution, with almost no overlap between the size of the minor and major workers. The major workers are approximately 8–10 mm (0.31–0.39 in) in length and the minors approximately half the length of the majors. Major workers forage, defend, maintain, and expand the colony whereas minor workers tend to stay within the nests where they care for the brood and 'milk' scale insects in or close to the nests.

<span class="mw-page-title-main">Carpenter ant</span> Genus of ants (Camponotus spp.)

Carpenter ants are large ants indigenous to many forested parts of the world.

<span class="mw-page-title-main">Banded sugar ant</span> Species of carpenter ant (Camponotus consobrinus)

The banded sugar ant, also known as the sugar ant, is a species of ant native to Australia. A member of the genus Camponotus in the subfamily Formicinae, it was described by German entomologist Wilhelm Ferdinand Erichson in 1842. Its common name refers to the ant's liking for sugar and sweet food, as well as the distinctive orange-brown band that wraps around its gaster.

<span class="mw-page-title-main">Meat ant</span> Common Australian ant

The meat ant, also known as the gravel ant or southern meat ant, is a species of ant endemic to Australia. A member of the genus Iridomyrmex in the subfamily Dolichoderinae, it was described by British entomologist Frederick Smith in 1858. The meat ant is associated with many common names due to its appearance, nest-building behaviour and abundance, of which its specific name, purpureus, refers to its coloured appearance. It is among the best-known species of ant found throughout Australia; it occurs in almost all states and territories except for Tasmania. Its enormous distribution, aggression and ecological importance have made this ant a dominant species.

<span class="mw-page-title-main">Black carpenter ant</span> Species of American ant (Camponotus pennsylvanicus)

The black carpenter ant is one of the largest and most common species of carpenter ant native to the central and eastern United States as well as eastern Canada.

<i>Myrmecia nigriceps</i> Endemic ant species from Australia

Myrmecia nigriceps, also known as the black-headed bull ant, is a species of ant endemic to Australia. A member of the genus Myrmecia in the subfamily Myrmeciinae, it was first described by Austrian entomologist Gustav Mayr in 1862. These ants are large, varying from 19 to 23 millimetres in length. However, colonies contain workers that are much smaller, usually half the size of normal workers. The queens are the largest while the males are the smallest, which can be easily identified due to their small mandibles.

<span class="mw-page-title-main">Green-head ant</span> Species of ant

The green-head ant is a species of ant that is endemic to Australia. It was described by British entomologist Frederick Smith in 1858 as a member of the genus Rhytidoponera in the subfamily Ectatomminae. These ants measure between 5 and 7 mm. The queens and workers look similar, differing only in size, with the males being the smallest. They are well known for their distinctive metallic appearance, which varies from green to purple or even reddish-violet. Among the most widespread of all insects in Australia, green-head ants are found in almost every Australian state, but are absent in Tasmania. They have also been introduced in New Zealand, where several populations have been established.

<i>Myrmeciites</i> Extinct genus of ants

Myrmeciites is an extinct form genus of bulldog ants in the subfamily Myrmeciinae of the family Formicidae, which contains three described species and two fossils not placed beyond the genus level. Described in 2006 from Ypresian stage deposits, all three of the described species and one unplaced fossil are from British Columbia, Canada, while the second unplaced fossil is from Washington State, USA. These ants were large, with the largest specimens collected reaching 3 centimetres (1.2 in). The behaviour of these ants would have been similar to extant Myrmeciinae ants, such as solitary foraging, nesting either in the soil or trees, and leaving no pheromone trail to food sources. Due to the poor preservation of these ants, their phylogenetic position among Myrmeciinae is unclear, and no type species has been designated. These ants are classified as incertae sedis in Myrmeciinae, but some writers have classified it as incertae sedis within the insect order Hymenoptera. This reclassification, however, has not been accepted; instead, Myrmeciites remains in Myrmeciinae.

<i>Leptothorax acervorum</i> Species of ant

Leptothorax acervorum is a small brown to yellow ant in the subfamily Myrmicinae. It was first described by Johan Christian Fabricius in 1793. L. acervorum is vastly distributed across the globe, most commonly found in the coniferous forests of Central, Western and Northern Europe. The morphology of L. acervorum is extremely similar to that of other Leptothorax ants. The difference arises in the two-toned appearance of L. acervorum, with the head and metasoma being darker than the mesosoma segment of the body, and hair across its body. Following Bergmann's rule—unusually, for ectothermic animals—body size increases with latitude.

<i>Formica pallidefulva</i> Species of ant

Formica pallidefulva is a species of ant found in North America. It is a red to dark brown ant with a shiny body, and varies in shade across its range. Colonies of this ant are found in a variety of habitats, where they excavate underground nests with galleries and chambers. In some parts of its range, the nests may be raided by slave-making ants, most notably Formica pergandei and Polyergus montivagus.

<i>Myrmecia regularis</i> Species of ant endemic to Australia

Myrmecia regularis is a species of ant endemic to Australia. A member of the genus Myrmecia in the subfamily Myrmeciinae, it was first described by American entomologist Walter Cecil Crawley in 1925. These ants are medium to large in size, measuring 10 to 20 millimetres, and they are bright brownish-red in colour. Queens and workers share similar morphological features, but they can be distinguished by the noticeable size difference. Males also look similar, but collected specimens are too damaged to be examined properly.

<span class="mw-page-title-main">Ergatoid</span> Wingless reproductive ant or termite

An ergatoid is a permanently wingless reproductive adult ant or termite. The similar but somewhat ambiguous term ergatogyne refers to any intermediate form between workers and standard gynes. Ergatoid queens are distinct from other ergatogyne individuals in that they are morphologically consistent within a species and are always capable of mating, whereas inter caste individuals, another class of ergatogynes, often are not. Ergatoids can exhibit wide morphological differences between species, sometimes appearing almost identical to normal workers and other times being quite distinct from both workers and standard queens. In addition to morphological features, ergatoids among different species can exhibit a wide range of behaviors, with some ergatoids acting only as reproductives and others actively foraging. Ergatoid queens have developed among a large number of ant species, and their presence within colonies can often provide clues on the social structures of colonies and as to how new colonies are founded. Without wings, almost all species of ants that solely produce ergatoid queens establish new colonies by fission.

<i>Agelaia pallipes</i> Species of wasp

Agelaia pallipes is a species of social paper wasp found from Costa Rica to Brazil, Argentina and Paraguay. A. pallipes is ground-nesting and is one of the most aggressive wasps in South America. This species is a predator of other insects, including flies, moths, and ground crickets, as well as baby birds.

<i>Camponotus floridanus</i> Species of ant known as the Florida carpenter ant

Camponotus floridanus, or Florida carpenter ant, is a species of ant in the genus Camponotus. First described as Formica floridana by Buckley in 1866, the species was moved to Camponotus by Mayr in 1886. The ant is widespread in Florida and occurs as far north as North Carolina and as far west as Mississippi.

<i>Camponotus socius</i> Species of ant

Camponotus socius, the sandhill carpenter ant, is a large species of ant in the genus Camponotus. It was first described by Julius Roger (1863), based on specimens from Brazil - however these can be considered highly dubious as the location where the type specimens were collected (Amazonas) does not fit the known ecology of the species within North America, where it exhibits traits typical of a native species. It is well adapted to the sandy soils of xeric woodlands found within the coastal plains of the southeastern United States. Its range includes the US states of Georgia, Alabama, Florida, North and South Carolina and Mississippi.

<span class="mw-page-title-main">Black-headed sugar ant</span> Species of carpenter ant (Camponotus nigriceps)

The black-headed sugar ant, also known as the brown sugar ant, is a species of Formicinae ant endemic to Australia. Found throughout most states, the species is a member of the genus Camponotus, a cosmopolitan genus of ants commonly known as carpenter ants. It was formally described and named by British entomologist Frederick Smith in 1858. These ants are characterised by their black head, reddish-brown mesosoma and black gaster, which can change in colour.

This is a glossary of terms used in the descriptions of ants.

<i>Bradoponera</i> Extinct genus of ants

Bradoponera is an extinct genus of ant in the Formicidae subfamily Proceratiinae, and is one of four genera of the subfamily. The genus contains four described species Bradoponera electrina, Bradoponera meieri, Bradoponera similis, and Bradoponera wunderlichi. The species are known from several Middle Eocene amber fossils which were found in Europe.

<i>Camponotus ligniperda</i> Brown-black carpenter ant

Camponotus ligniperda, the brown-black carpenter ant, is a common species of carpenter ant distributed widely throughout Europe. Found in a variety of woodland habitats, they commonly nest on the ground in dry tree stumps, dead fallen trees, or beneath stones and wooden logs that are partially buried. C. ligniperda is an ecologically dominant species wherever it is found due to both its large size and particularly aggressive nature.

References

  1. 1 2 "Camponotus herculeanus (carpenter ants)". AntKeepers. Retrieved 7 April 2021.
  2. Bolton, B. (2015). "Camponotus herculeanus". AntCat. Retrieved 31 January 2015.
  3. Linnaeus, Carolus (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata (in Latin). Holmiae. (Laurentii Salvii). p. 579.
  4. Mayr, G. (1861). Die europäischen Formiciden. Nach der analytischen Methode bearbeitet. Wien: C. Gerolds Sohn, 80 pp.
  5. 1 2 3 4 5 "Camponotus herculeanus". AntWiki. Retrieved 30 March 2021.
  6. Vander Meer, Robert K (23 April 2019). Applied Myrmecology: A World Perspective. CRC Press. pp. 274–277. ISBN   978-0-429-72218-9.
  7. Smith, Falconer (1944). "Nutritional requirements of Camponotus ants". Annals of the Entomological Society of America. 37 (4): 401–408. doi:10.1093/aesa/37.4.401.