Period | Upper Paleolithic, Mesolithic |
---|---|
Dates | 13,000–6,000 BC |
Caucasus hunter-gatherer (CHG), also called Satsurblia cluster, [1] is an anatomically modern human genetic lineage, first identified in a 2015 study, [2] [3] based on the population genetics of several modern Western Eurasian (European, Caucasian and Near Eastern) populations. [4] [5]
The CHG lineage is suggested to have diverged from the ancestor of Western Hunter-Gatherers (WHGs) probably during the Last Glacial Maximum (sometimes between 45,000 to 26,000 years ago). [6] They further separated from the "Anatolian Hunter Gatherer" (AHG) lineage later, suggested to around 25,000 years ago during the late LGM period. [3] [7] The Caucasus hunter-gatherers managed to survive in isolation since the late LGM period as a distinct population, and display high genetic affinities to Mesolithic and Neolithic populations on the Iranian plateau, such as Neolithic specimens found in Ganj Dareh. The CHG display higher genetic affinities to European and Anatolian groups than Iranian hunter-gatherers do, suggesting a possible cline and geneflow into the CHG and less into Mesolithic and Neolithic Iranian groups. [2] [8]
The Mesolithic/Neolithic Iranian lineage and the Caucasus hunter-gatherers are inferred to derive significant amounts of their ancestry from Basal Eurasian (c. 48%), with the remainder ancestry being closer to Ancient North Eurasians (ANE; c. 52%). The CHG displayed an additional ANE-like component (c. 10%) than the Neolithic Iranians do, suggesting they may have stood in continuous contact with Eastern Hunter-Gatherers to their North. The CHG also carry around 20% additional Paleolithic Caucasus/Anatolian ancestry. [2] [9] An alternative model without the need of significant amounts of ANE ancestry has been presented by Vallini et al. 2024, suggesting that the initial Iranian hunter-gatherer-like population which is basal to the CHG formed primarily from a deep Ancient West Eurasian lineage (WEC2, at least 50%), and from varying degrees of Ancient East Eurasian and Basal Eurasian components. The Ancient West Eurasian component associated with Iranian hunter-gatherers is inferred to have diverged from the West Eurasian Core lineage (represented by Kostenki-14; WEC), with the WEC2 component staying in the region of the Iranian Plateau, while the proper WEC component expanded into Europe. [10]
At the beginning of the Neolithic, at c. 8000 BC, they were probably distributed across western Iran and the Caucasus, [11] and people similar to northern Caucasus and Iranian plateau hunter-gatherers arrived before 6000 BC in Pakistan and north-west India. [12] A roughly equal merger between the CHG and Eastern Hunter-Gatherers in the Pontic–Caspian steppe resulted in the formation of the Western Steppe Herders (WSHs). The WSHs formed the Yamnaya culture and subsequently expanded massively throughout Europe during the Late Neolithic and Early Bronze Age circa 3000-2000 BC. [13]
Jones et al. (2015) analyzed genomes from males from western Georgia, in the Caucasus, from the Late Upper Palaeolithic (13,300 years old) and the Mesolithic (9,700 years old). These two males carried Y-DNA haplogroup: J* and J2a, later refined to J1-FT34521, and J2-Y12379*, and mitochondrial haplogroups of K3 and H13c, respectively. [14] Their genomes showed that a continued mixture of the Caucasians with Middle Eastern populations took place up to 25,000 years ago, when the coldest period in the last Ice Age started. [4]
CHG ancestry was also found in an Upper Palaeolithic specimen from Satsurblia cave (dated ca. 11000 BC), and in a Mesolithic one from Kotias Klde cave, in western Georgia (dated ca. 7700 BC). The Satsurblia individual is closest to modern populations from the South Caucasus. [2]
Margaryan et al. (2017) analysing South Caucasian ancient mitochondrial DNA found a rapid increase of the population at the end of the Last Glacial Maximum, about 18,000 years ago. The same study also found continuity in descent in the maternal line for 8,000 years. [15]
According to Narasimhan et al. (2019) Iranian farmer related people arrived before 6000 BCE in Pakistan and north-west India, before the advent of farming in northern India. They suggest the possibility that this "Iranian farmer–related ancestry [...] was [also] characteristic of northern Caucasus and Iranian plateau hunter-gatherers." [12]
The ancestry of the Yamnaya people can be mostly modelled as an admixture of Eastern Hunter-Gatherers (EHGs) and a Near Eastern component related to Caucasus hunter-gatherers, Iranian Chalcolithic people, or a genetically similar population. [17] [2] [18] [4] [note 1] Each of those two populations contributed about half the Yamnaya DNA. [19] [4] According to co-author Andrea Manica of the University of Cambridge:
The question of where the Yamnaya come from has been something of a mystery up to now […] we can now answer that, as we've found that their genetic make-up is a mix of Eastern European hunter-gatherers and a population from this pocket of Caucasus hunter-gatherers who weathered much of the last Ice Age in apparent isolation. [4]
According to Jones et al. (2015), Caucasus hunter-gatherers (CHG) "genomes significantly contributed to the Yamnaya steppe herders who migrated into Europe ~3,000 BCE, supporting a formative Caucasus influence on this important Early Bronze Age culture. CHG left their imprint on modern populations from the Caucasus and also Central and South Asia possibly correlating with the arrival of Indo-Aryan languages." [20]
Lazaridis et al. (2016) proposes a different people, likely from Iran, as the source for the Middle Eastern ancestry of the Yamnaya people, finding that "a population related to the people of the Iran Chalcolithic contributed ~43% of the ancestry of early Bronze Age populations of the steppe". [21] [note 2] That study asserts that these Iranian Chalcolithic people were a mixture of "the Neolithic people of western Iran, the Levant, and Caucasus Hunter Gatherers". [21]
Gallego-Llorente et al. (2016) conclude that Iranian populations are not a likelier source of the 'southern' component in the Yamnaya than Caucasus hunter-gatherers. [22]
Wang et al. (2018) analysed genetic data of the North Caucasus of fossils dated between the 4th and 1st millennia BC and found correlation with modern groups of the South Caucasus, concluding that "unlike today – the Caucasus acted as a bridge rather than an insurmountable barrier to human movement". [23]
While some argue that the Pre-Proto-Indo-European language may have originated among a CHG-rich population in Western Asia, [24] others, such as David W. Anthony, suggest that the Indo-European languages were initially spoken by EHGs living in Eastern Europe. [25]
Beyond contributing to the population of mainland Europe through Bronze Age pastoralists of the Yamnaya, CHG also appears to have arrived on its own in the Aegean without Eastern European hunter–gatherer (EHG) ancestry and provided approximately 9–32% of ancestry to the Minoans. The origin of this CHG component might have been Central Anatolia. [26]
The Proto-Indo-Europeans are a hypothetical prehistoric ethnolinguistic group of Eurasia who spoke Proto-Indo-European (PIE), the reconstructed common ancestor of the Indo-European language family.
The Corded Ware culture comprises a broad archaeological horizon of Europe between c. 3000 BC – 2350 BC, thus from the late Neolithic, through the Copper Age, and ending in the early Bronze Age. Corded Ware culture encompassed a vast area, from the contact zone between the Yamnaya culture and the Corded Ware culture in south Central Europe, to the Rhine on the west and the Volga in the east, occupying parts of Northern Europe, Central Europe and Eastern Europe. Early autosomal genetic studies suggested that the Corded Ware culture originated from the westward migration of Yamnaya-related people from the steppe-forest zone into the territory of late Neolithic European cultures; however, paternal DNA evidence fails to support this hypothesis, and it is now proposed that the Corded Ware culture evolved in parallel with the Yamnaya, with no evidence of direct male-line descent between them.
The Yamnaya culture or the Yamna culture, also known as the Pit Grave culture or Ochre Grave culture, is a late Copper Age to early Bronze Age archaeological culture of the region between the Southern Bug, Dniester, and Ural rivers, dating to 3300–2600 BCE. It was discovered by Vasily Gorodtsov following his archaeological excavations near the Donets River in 1901–1903. Its name derives from its characteristic burial tradition: Я́мная is a Russian adjective that means 'related to pits ', as these people used to bury their dead in tumuli (kurgans) containing simple pit chambers.
The Srubnaya culture, also known as Timber-grave culture, was a Late Bronze Age 1900–1200 BC culture in the eastern part of the Pontic–Caspian steppe. It is a successor of the Yamna culture, the Catacomb culture and the Poltavka culture. It is co-ordinate and probably closely related to the Andronovo culture, its eastern neighbor. Whether the Srubnaya culture originated in the east, west, or was a local development, is disputed among archaeologists.
The Sredny Stog culture is a pre-Kurgan archaeological culture from the 5th–4th millennia BC. It is named after the Dnieper river islet of today's Serednii Stih, Ukraine, where it was first located.
The Catacomb culture was a Bronze Age culture which flourished on the Pontic steppe in 2,500–1,950 BC.
The Khvalynsk culture is a Middle Copper Age Eneolithic culture of the middle Volga region. It takes its name from Khvalynsk in Saratov Oblast. The Khvalynsk culture is found from the Samara Bend in the north to the North Caucasus in the south, from the Sea of Azov in the west to the Ural River in the east. It was preceded by the Early Eneolithic Samara culture.
The Dnieper–Donets culture complex (DDCC) is a Mesolithic and later Neolithic archaeological culture found north of the Black Sea and dating to ca. 5000-4200 BC. It has many parallels with the Samara culture, and was succeeded by the Sredny Stog culture.
The genetic history of Europe includes information around the formation, ethnogenesis, and other DNA-specific information about populations indigenous, or living in Europe.
The Armenian hypothesis, also known as the Near Eastern model, is a theory of the Proto-Indo-European homeland, initially proposed by linguists Tamaz V. Gamkrelidze and Vyacheslav Ivanov in the early 1980s, which suggests that the Proto-Indo-European language was spoken during the 5th–4th millennia BC in "eastern Anatolia, the southern Caucasus, and northern Mesopotamia".
The Proto-Indo-European homeland was the prehistoric linguistic homeland of the Proto-Indo-European language (PIE). From this region, its speakers migrated east and west, and went on to form the proto-communities of the different branches of the Indo-European language family.
The peopling of India refers to the migration of Homo sapiens into the Indian subcontinent. Anatomically modern humans settled India in multiple waves of early migrations, over tens of millennia. The first migrants came with the Coastal Migration/Southern Dispersal 65,000 years ago, whereafter complex migrations within South and Southeast Asia took place. West-Asian (Iranian) hunter-gatherers migrated to South Asia after the Last Glacial Period but before the onset of farming. Together with ancient South Asian hunter-gatherers they formed the population of the Indus Valley Civilisation (IVC).
The genetic history of Italy is greatly influenced by geography and history. The ancestors of Italians were mostly Indo-European speaking peoples and pre-Indo-European speakers. During the Roman empire, the Italian peninsula attracted people from various regions of the Mediterranean basin, including Southern Europe, North Africa and the Middle East. Based on DNA analysis, there is evidence of ancient regional genetic substructure and continuity within modern Italy dating to the pre-Roman and Roman periods.
The Indo-European migrations are hypothesized migrations of Proto-Indo-European language (PIE) speakers, and subsequent migrations of people speaking derived Indo-European languages, which took place approx. 4000 to 1000 BCE, potentially explaining how these languages came to be spoken across a large area of Eurasia, spanning from the Indian subcontinent and Iranian plateau to Atlantic Europe, in a process of cultural diffusion.
In archaeogenetics, the term Ancient North Eurasian (ANE) is the name given to an ancestral component that represents the lineage of the people of the Mal'ta–Buret' culture (c. 24,000 BP) and populations closely related to them, such as the Upper Paleolithic individuals from Afontova Gora in Siberia. Genetic studies also revealed that the ANE are closely related to the remains of the preceding Yana Culture (c. 32,000 BP), which were dubbed as 'Ancient North Siberians' (ANS), and which either are directly ancestral to the ANE, or both being closely related sister lineages, sharing a common origin from the merger of an 'Early West Eurasian' hunter-gatherer lineage (represented by Kostenki-14, c. 38,000 BP) and an 'Early East Eurasian' population (represented by the Tianyuan man, c. 40,000 BP). The Ancient North Eurasians derive around 2/3 from an Early West Eurasian lineage and around 1/3 of their ancestry from an Early East Eurasian lineage.
Early European Farmers (EEF), First European Farmers, Neolithic European Farmers, Ancient Aegean Farmers, or Anatolian Neolithic Farmers are names used to describe a distinct group of early Neolithic farmers who brought agriculture to Europe and Northwest Africa (Maghreb). Although the spread of agriculture from the Middle East to Europe has long been recognised through archaeology, it is only recent advances in archaeogenetics that have confirmed that this spread was strongly correlated with a migration of these farmers, and was not just a cultural exchange.
In archaeogenetics, the term Western Hunter-Gatherer (WHG), West European Hunter-Gatherer, Western European Hunter-Gatherer, Villabruna cluster, or Oberkassel cluster is the name given to a distinct ancestral component of modern Europeans, representing descent from a population of Mesolithic hunter-gatherers who scattered over Western, Southern and Central Europe, from the British Isles in the west to the Carpathians in the east, following the retreat of the ice sheet of the Last Glacial Maximum.
In archaeogenetics, the term Eastern Hunter-Gatherer (EHG), sometimes East European Hunter-Gatherer, or Eastern European Hunter-Gatherer is the name given to a distinct ancestral component that represents Mesolithic hunter-gatherers of Eastern Europe.
Basal Eurasian is a proposed lineage of anatomically modern humans with reduced, or zero, archaic hominin (Neanderthal) admixture compared to other ancient non-Africans. Basal Eurasians represent a sister lineage to other Eurasians and may have originated from the Southern Middle East, specifically the Arabian peninsula, or North Africa, and are said to have contributed ancestry to various West Eurasian, South Asian, and Central Asian as well as African groups. This Basal Eurasian component is also proposed to explain the lower archaic admixture among modern West Eurasians compared to with East Eurasians, although alternatives without the need of such Basal admixture exist as well.
In archaeogenetics, the term Western Steppe Herders (WSH), or Western Steppe Pastoralists, is the name given to a distinct ancestral component first identified in individuals from the Chalcolithic steppe around the turn of the 5th millennium BC, subsequently detected in several genetically similar or directly related ancient populations including the Khvalynsk, Sredny Stog, and Yamnaya cultures, and found in substantial levels in contemporary European, West Asian and South Asian populations. This ancestry is often referred to as Yamnaya ancestry, Yamnaya-related ancestry, Steppe ancestry or Steppe-related ancestry.
By reading the DNA, the researchers were able to show that the lineage of this fourth Caucasus hunter-gatherer strand diverged from the western hunter-gatherers just after the expansion of anatomically modern humans into Europe from Africa.