Peopling of China

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A population genomic PCA graph, showing the substructure of East Asian populations, including Han Chinese (2019) East Asian PCA (including Jomon samples).png
A population genomic PCA graph, showing the substructure of East Asian populations, including Han Chinese (2019)

In the course of the peopling of the World by Homo sapiens , East Asia was reached about 50,000 years ago (50 kya). The "recent African origin" lineage of 70 kya diverged into identifiable East Eurasian and West Eurasian lineages by about 50 kya. [2] [3] The East Eurasian ancestors of East Asians used a southern route to reach South and Southeast Asia, along which they rapidly diverged into the ancestors of Indigenous South Asians (AASI), Papuans, East Asians and Andamanese peoples (such as the Onge). [4] This early East Asian lineage diverged further during the Last Glacial Maximum, contributing outgoing from Mainland Southeast Asia significantly to the peopling of the Americas via Beringia about 25 kya. [5] After the last ice age China became cut off from neighboring island groups. The previous phenotypes of early East Asians became either replaced or prevailed among more geographically distant groups. [6] [7]

Contents

Genetic history

Overview

A "Basal-East Asian population" referred to as the East- and Southeast Asian lineage (ESEA); is ancestral to modern East Asians, Southeast Asians, Polynesians, and Siberians, but also ancestral to the Hoabinhian hunter-gatherers of Southeast Asia and the ~40,000 year old Tianyuan lineage found in Northern China. The ESEA lineage descend from an earlier "eastern non-African" (ENA) or "Ancient East Eurasian" meta-population, which used a single southern route to reach South, Southeast Asia, and Oceania, and along which they rapidly diverged into the ancestors of Ancient South Asians (AASI), East/Southeast Asians (ESEA), as well as Australasians. The early ESEA lineage is inferred to have than diverged into the Hoabinhian, the Tianyuan, and Ancient East Asian lineages, and expanded northward. There is "a strong correlation with latitude, with diversity decreasing from south to north". [8] [4]

Genetic studies of Shaanxi and Liaoning

One study showed some Han Chinese individuals in Shaanxi carry maternal haplogroup U. [9] Other mtDNAs that some Han Chinese have are W6 and H. Some Han Chinese also carried paternal haplogroup R1a1. [10]

Several studies reveal minor West Eurasian-derived admixture among Shaanxi Han Chinese, especially those living in Guanzhong and Shaanbei (2–5%), [10] [11] and Liaoning Chinese (~2%). [12] Ancient North Eurasian admixture is more dominant among Shaanxi Han Chinese compared to other Han subgroups. [10]

Archaeogenetic studies in Guangxi

Wang et al. (2021) found that the ancestries of individuals in Guangxi from between 9,000–6,000 BP can be modeled as mixtures of Upper Paleolithic source populations from both Guangxi and Fujian. During the early and late Neolithic, migration of Austronesians from the Fujian region's Neolithic Austronesians significantly influenced the genetic profile of the Guangxi region but did not completely replace the earlier resident hunter-gatherer ancestry that was local to Guangxi. Neolithic Fujian is genetically unrelated to Modern Fujian. The main archaic individuals that were analyzed include the following.

Huang et al. (2022) associate Dushan and Baojianshan-related ancestry with the first Neolithic farmers in Mainland Southeast Asia (MSEA), i.e. late Neolithic farmers who expanded from southern China into MSEA. Among present-day populations, they found that: [14]

Guangxi inhabitants from 1,500–500 BP contributed to the ancestries of present-day Kra-Dai and Hmong-Mien-speaking populations of southern China. They can be modeled as a mixture of 58.2%–90.6% Dushan-related (or Qihe-3-related) ancestry and 9.4%–41.8% northern East Asian-related ancestry, originating from Shandong about 9,500–7,700 years ago. [15]

Paternal lineages

A PCA graph illustrating the genetic makeup of Han Chinese and other Asian populations. 1 asia.png
A PCA graph illustrating the genetic makeup of Han Chinese and other Asian populations.
Hypothetical migration patterns of paternal human lineages Migraciones humanas en haplogrupos de ADN-Y.PNG
Hypothetical migration patterns of paternal human lineages
Haplogroup O, also known as O-M175, is primarily found among populations in Southeast Asia and China. Haplogrupo O (ADN-Y).PNG
Haplogroup O, also known as O-M175, is primarily found among populations in Southeast Asia and China.

Looking at Y-DNA studies, it would seem that East Asian paternal lineages expanded in Asia approximately 50,000 years ago. People bearing genetic markers ancestral such as C, D, N, and O, as well as P (specifically Q), came through the Himalayan mountain range and proceeded to Southeast Asia. [17] [18] Haplogroup C moved to East Asia and Australia, with at least two subclades of the major East Asian branch migrating into the Americas, and with members of Haplogroup C-M38 spreading throughout Wallacea, New Guinea, Melanesia, and Polynesia. Another group of peoples, bearing the Y-DNA Haplogroup D, has left descendants mostly in the Andaman Islands, Tibet, and Japan. [19] [20] Haplogroup Q, believed to have arisen in Central Asia or Southern Siberia approximately 17,000 to 22,000 years ago, went north to populate Northern Siberia and the Americas. Some northern Chinese have this genetic marker. Haplogroups N and O, originated in Southern China and by 10,000 years ago went on to populate first Southeast Asia and then from Southeast Asia, left for East Asia. Roughly 12,000 years ago, during the Neolithic period, farmers settled along the Yellow River. Alongside various other lineages including O2‐M122, they initiated the development of agriculture. About 6000 years ago, ancestors of the Tibetans split off from this parent group. About 5,000 years ago, Neolithic Yellow River farmers experienced rapid expansion, with notable gene flow into surrounding populations. [21] This corresponds to the late period (2600-2000 BC) of the Longshan culture in the middle Yellow River area. As the Neolithic population in China reached its peak, the number of settlements increased. In some locations, such as the basin of the Fen River in southern Shanxi, the Yellow River in western Henan (confined by the Zhongtiao Mountains and Xiao Mountains), and the coastal Rizhao plain of southeast Shandong, a few very large (over 200 ha) centers developed. In more open areas, such as the rest of Shandong, the Central Plain (in Henan) and the Wei River basin in Shaanxi, local centers were more numerous, smaller (generally 20 to 60 ha) and fairly evenly spaced. Walls of rammed earth have been found in 20 towns in Shandong, nine in the Central Plain, and one (Taosi) in southern Shanxi, suggesting conflict between polities in these areas.

The expansion and rise of these various settlements could be due to the impetus of the collective benefit of the construction of irrigation works in the late Neolithic:

"Most of the labor to dike and drain an area is associated with digging a ditch and sidecasting the soil to make an earthen dike. To make the culvert and tide gate you can use an old worn-out canoe for the pipe .... and use just about any good-sized flat stones you can get your hands on. If you have twice as many people making your dike, you can make twice as many linear feet of dike. Doubling the perimeter of a square dike results in quadrupling the area within the dike."

The agricultural surplus would have allowed for a rapidly expanding population, which would provide more labor for irrigation. By the time of the establishment of the Xia and Shang dynasties, population estimates were at approximately 13 million people. [22]

Studies of DNA remnants from the Central Plains area of China 3000 years ago show close affinity between that population and those of Northern Han today in both the Y-DNA and mtDNA. Both northern and southern Han show similar Y-DNA genetic structure. [23]

Y-chromosome haplogroup O2-M122 is a common DNA marker in Han Chinese, as it appeared in China in prehistoric times. It is found in more than 50% of Chinese males, with frequencies tending to be high toward the east of the country (30/101 = 29.7% Guangxi Pinghua Han, [24] 13/40 = 32.5% Guangdong Han, [25] 11/30 = 36.7% Lanzhou Han, [26] 26/60 = 43.3% Yunnan Han, [27] 251/565 = 44.4% Zhaotong Han, [28] 15/32 = 46.9% Yili Han, [26] 23/49 = 46.9% Lanzhou Han, [29] [30] 32/65 = 49.2% South China Han, [31] 18/35 = 51.4% Meixian Han, [26] 22/42 = 52.4% Northern Han, [32] 43/82 = 52.4% Northern Han, [33] 18/34 = 52.9% Chengdu Han, [26] 154/280 = 55.0% Southern Han, [33] 27/49 = 55.1% Northern Han, [34] 73/129 = 56.6% North China Han, [31] 49/84 = 58.3% Taiwan Han, [25] 35/60 = 58.3% Taiwan Minnan, [35] 99/167 = 59.3% East China Han, [31] 33/55 = 60.0% Fujian Han, [35] 157/258 = 60.9% Taiwan Han, [35] 13/21 = 61.9% Taiwan Han, [34] 189/305 = 62.0% Zibo Han, [28] 23/35 = 65.7% Harbin Han, [26] 29/44 = 65.9% Northern Han, [25] 23/34 = 67.6% Taiwan Hakka, [35] 35/51 = 68.6% Beijing Han [27] ). [36] [37]

During the Zhou dynasty, or earlier, peoples with haplogroup Q-M120 likewise also contributed to the ethnogenesis of Han Chinese people. This haplogroup is implied to be widespread in the Eurasian steppe and north Asia since it is found among Cimmerians in Moldova and Bronze Age natives of Khövsgöl. But it is currently near-absent in these regions except for East Asia. In modern China, haplogroup Q-M120 can be found in the northern and eastern regions. [38]

Han Chinese are genetically distinguishable from Yamato Japanese and Koreans, and internally the different Han Chinese subgroups are genetically closer to each other than any of them are to Koreans and Japanese. However, some Southern Han Chinese, such as Guangxi Han, are genetically closer to Vietnamese and Dai people than Northern Han. But meanwhile, when compared to Europeans genetics, the Han Chinese, Southeast Asian, Japanese and Koreans are closer to each other than Europeans and South Asians. Genealogical research has indicated extremely similar genetic profiles of a less than 1% total variation in spectrum between these three groups. [39] Some Southern Han Chinese and Northern Han Chinese are closest to each other and show the smallest differences when they are compared to other Asians. Vietnamese Ho Chi Minh City Kinh are close to Xishuangbanna's Dai ethnic minority and Guangdong Han Chinese. [40] Koreans are also relatively close to northern Han Chinese. Japanese are more genetically distant from Koreans than Koreans are from northern Han Chinese. However Buryat Mongols and Qinghai Mongols are further from each other than Japanese and Korean despite both being Mongols. [41] Comparisons between the Y chromosome SNP and MtDNA of modern Northern Han Chinese and 3,000 year old Hengbei ancient samples from China's Central Plains show they are extremely similar to each other and show continuity between ancient Chinese of Hengbei and current Northern Han Chinese while Southern Han Chinese were different from the people of Hengbei. This showed that already 3,000 years ago the current northern Han Chinese genetic structure was already formed. [23]

Maternal lineages

The mitochondrial-DNA haplogroups of the Han Chinese can be classified into the northern East Asian-dominating haplogroups, including A, C, D, G, M8, M9, and Z, and the southern East Asian-dominating haplogroups, including B, F, M7, N*, and R. [42]

These haplogroups account for 52.7% and 33.85% of those in the Northern Han, respectively.

Haplogroup mtDNA D descend from Dravidian Haplogroup M mtdna is the modal mtDNA haplogroup among northern East Asians. Among these haplogroups, D, B, F, and A were predominant in the Northern Han, with frequencies of 25.77%, 11.54%, 11.54%, and 8.08%, respectively.

However, in the Southern Han, the northern and southern East Asian-dominating mtDNA haplogroups accounted for 35.62% and 51.91%, respectively. The frequencies of haplogroups D, B, F, and A reached 15.68%, 20.85%, 16.29%, and 5.63%, respectively. [43] [44] [45] [46] [47]

Climate history

During the Last Glacial Maximum, 29,000 to 18,000 years ago, northern China was a treeless steppe with areas of permafrost and southern China lost much of its forest cover. The sea level was much lower. Borneo, Indonesia, the Philippines, and the Japanese archipelago may have been accessible by land. With the end of the last ice age, a period of warming occurred lasting from 18,000 to 10,000 years ago. The oceans rose and inundated vast regions leaving little trace of coastal settlements used by these people. We know little about their languages. Their cultures are likely to have been diverse. There are many limestone cave sites in southern China which show human settlements. There is evidence of pottery making. The inhabitants had bone tools, fished, and hunted pigs and deer. [48]

It is believed that the climate in southern China was warmer and wetter south of the Qinling mountains; elephants are known to have inhabited the Yangtze river region. The climate in Northeast China north of present-day Beijing was characterized as a cold steppe environment during this period. The presence of woolly mammoth is well documented. [49]

The climate was also much warmer between 8,000 and 3,500 years ago. In the Shandong region, excavations have found the bones of alligators and elephants.

The development of agriculture about 10,000 years ago, with the domestication of millet in the Yellow River valley region and rice in the Yangtze River valley, may have been associated with accelerated growth in the number and size of settlements and the intensified development of local cultures and languages. [50]

Settlement patterns

Early settlements in the Chinese Upper Paleolithic were either hunter-gatherer societies, or marine environment based societies characterized by shell middens. [51] Relatively speaking the land was sparsely populated, as the peoples followed the coastal regions and the river valleys.

Neolithic settlements have been found from Liaoning province in the northeast to the Chengdu region in the southwest; from Gansu province in the northwest to sites in Fujian in the southeast. The settlement pattern in the Tibetan region is still unclear as there is debate as to whether there was a pre-Neolithic population movement into the region. [52]

Holocene China

The origin of Sino-Tibetan is uncertain. It dates to the Neolithic, or just before the Neolithic. There are various proposals. Van Driem (2005) proposes that Sino-Tibetan originated in the Sichuan Basin before 7000 BC, with an early migration into northeast India, and a later migration north of the predecessors of Chinese and Tibetic.[ citation needed ] Matisoff (1991) places its origins in the eastern part of the Tibetan plateau around 4000 BC, with the various groups migrating out along the Yellow, Yangtze, Mekong, Salween and Brahmaputra rivers.[ citation needed ] Blench and Post (2014) have proposed that the Sino-Tibetan homeland was in northeast India, the area of greatest diversity, around 7000 BC.[ citation needed ] Blench (2009) proposes that the earliest speakers of Sino-Tibetan were not farmers, since agriculture cannot be reconstructed for Proto-Sino-Tibetan. Rather, early Sino-Tibetan speakers were highly diverse hunter-gatherers and foragers. [53] More recent study published in the Proceedings of the National Academy of Sciences of the United States of America point to the origin of the Sino-Tibetan in northern China in the Neolithic Cishan culture and Yangshao culture. [54]

The Austroasiatic languages (including Vietnamese, Khmer, etc.) likely originated from Southern China and would have entered Southeast Asia via Yunnan and the Mekong. Likewise, the Miao, Thai, Burmese, and Tibetan languages evolved in China. It is even accepted that proto-Austronesian speakers originated from China, migrated to Taiwan (Formosa), and then to the islands of the Pacific, especially since Taiwan is the place exhibiting the most diversity in Austronesian languages (see Formosan languages). By 3,000 years ago, the sophistication of some of the cultures neighboring that of the Han Chinese can be seen in the bronze artifacts of the Sanxingdui (Sichuan), Ban Chiang (Thailand) and Dong Son (Vietnam). The domestication of the horse 6,000 years ago in the Eurasian steppes led to cultures that mixed animal husbandry and agriculture. Indo-European speakers are known to have reached the Xinjiang region 4,000 years ago (see Tocharians).

Because the East Asian landmass was initially populated from the south, and the historical record shows the Han Chinese migrating to the south and southwest, the genetic relationship among all the peoples of China remains a challenging study. [55]

See also

Related Research Articles

<span class="mw-page-title-main">Genetics and archaeogenetics of South Asia</span> Biological field of study

Genetics and archaeogenetics of South Asia is the study of the genetics and archaeogenetics of the ethnic groups of South Asia. It aims at uncovering these groups' genetic histories. The geographic position of the Indian subcontinent makes its biodiversity important for the study of the early dispersal of anatomically modern humans across Asia.

<span class="mw-page-title-main">Jōmon people</span> Early inhabitants of prehistoric Japan

Jōmon people is the generic name of the indigenous hunter-gatherer population that lived in the Japanese archipelago during the Jōmon period. They were united through a common Jōmon culture, which reached a considerable degree of sedentism and cultural complexity.

<span class="mw-page-title-main">Haplogroup M (mtDNA)</span> Widespread human mitochondrial DNA grouping indicating common ancestry

Haplogroup M is a human mitochondrial DNA (mtDNA) haplogroup. An enormous haplogroup spanning all the continents, the macro-haplogroup M, like its sibling the macro-haplogroup N, is a descendant of the haplogroup L3.

<span class="mw-page-title-main">Haplogroup F (mtDNA)</span> Human mitochondrial DNA haplogroup

Haplogroup F is a human mitochondrial DNA (mtDNA) haplogroup. The clade is most common in East Asia and Southeast Asia. It has not been found among Native Americans.

<span class="mw-page-title-main">Haplogroup C (mtDNA)</span> Human mitochondrial DNA haplogroup

In human mitochondrial genetics, Haplogroup C is a human mitochondrial DNA (mtDNA) haplogroup.

Haplogroup D1 or D-M174 is a subclade of haplogroup D-CTS3946. This male haplogroup is found primarily in East Asia, Magar-ethnic Nepal and the Andaman Islands. It is also found regularly with lower frequency in Central Asia, Siberia and Mainland Southeast Asia, and, more rarely, in Europe and the Middle East.

<span class="mw-page-title-main">Haplogroup F-M89</span> Human Y chromosome DNA grouping indicating common ancestry

Haplogroup F, also known as F-M89 and previously as Haplogroup FT, is a very common Y-chromosome haplogroup. The clade and its subclades constitute over 95% of paternal lineages outside of Africa.

<span class="mw-page-title-main">Haplogroup O-M175</span> Haplogroup O. Human Y chromosome DNA grouping common in Asia.

Haplogroup O, also known as O-M175, is a human Y-chromosome DNA haplogroup. It is primarily found among populations in Southeast Asia and East Asia. It also is found in various percentages of populations of the Russian Far East, South Asia, Central Asia, Caucasus, Crimea, Ukraine, Iran, Oceania, Madagascar and the Comoros. Haplogroup O is a primary descendant of haplogroup NO-M214.

Haplogroup O-M122 is an Eastern Eurasian Y-chromosome haplogroup. The lineage ranges across Southeast Asia and East Asia, where it dominates the paternal lineages with extremely high frequencies. It is also significantly present in Central Asia, especially among the Naiman tribe of Kazakhs.

<span class="mw-page-title-main">Tungusic peoples</span> Ethnolinguistic group

Tungusic peoples are an ethnolinguistic group formed by the speakers of Tungusic languages. They are native to Siberia, China, and Mongolia.

In human genetics, Haplogroup O-M119 is a Y-chromosome DNA haplogroup. Haplogroup O-M119 is a descendant branch of haplogroup O-F265 also known as O1a, one of two extant primary subclades of Haplogroup O-M175. The same clade previously has been labeled as O-MSY2.2.

<span class="mw-page-title-main">Haplogroup C-M217</span> Human Y-chromosome DNA haplogroup

Haplogroup C-M217, also known as C2, is a Y-chromosome DNA haplogroup. It is the most frequently occurring branch of the wider Haplogroup C (M130). It is found mostly in Central Asia, Eastern Siberia and significant frequencies in parts of East Asia and Southeast Asia including some populations in the Caucasus, Middle East, South Asia, East Europe. It is found in a much more widespread area with a low frequency of less than 2%.

<span class="mw-page-title-main">Haplogroup D-M15</span> Human Y-chromosome DNA haplogroup

Its phylogenetically closest relatives are found among the peoples of Japan, Central Asia, and the Andaman Islands in the Bay of Bengal. It is more distantly related to the Haplogroup D*, whose sub-clades are common throughout Asia.

In human mitochondrial genetics, Haplogroup G is a human mitochondrial DNA (mtDNA) haplogroup.

Tianyuan man are the remains of one of the earliest modern humans to inhabit East Asia. In 2007, researchers found 34 bone fragments belonging to a single individual at the Tianyuan Cave near Beijing, China. Radiocarbon dating shows the bones to be between 42,000 and 39,000 years old, which may be slightly younger than the only other finds of bones of a similar age at the Niah Caves in Sarawak on the South-east Asian island of Borneo.

<span class="mw-page-title-main">Y-DNA haplogroups in populations of East and Southeast Asia</span>

The tables below provide statistics on the human Y-chromosome DNA haplogroups most commonly found among ethnolinguistic groups and populations from East and South-East Asia.

<span class="mw-page-title-main">Liao civilization</span> Ancient Northeast Asian civilization

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<span class="mw-page-title-main">Haplogroup O-M117</span> Descendant branch of haplogroup O2a (formerly O3a)

Haplogroup O2a2b1a1-M117 or Haplogroup O2a2b1a1-M117 is a subclade of O2a2b1-M134 that occurs frequently in China and in neighboring countries like Bhutan, Nepal, and Korea, also found among Sino-Tibetan language speaking people.

Various theories have been proposed by scholars over time in regards to the genetic makeup and population history of East Asian peoples and their connection to genetically related populations such as Southeast Asians and North Asians, as well as Oceanians, and partly, Central Asians, South Asians, and Native Americans. These scholars have come to certain conclusions for the makeup of East Asians, collectively referred to as "East Eurasians" in population genomics, both through genomic research and trait-based analysis.

<span class="mw-page-title-main">Ancient Southern East Asian</span> Archaeogenetic name for an ancestral genetic component

In archaeogenetics, Ancient Southern East Asian (ASEA), also known as Southern East Asian (sEA), is an ancestral lineage that is represented by individuals from Qihe Cave in Fujian and Liangdao Island in the Taiwan Strait as well as Guangxi. Ancient Southern East Asian ancestry significantly contributed to the genetic makeup of modern populations in East Asia, Mainland Southeast Asia, Insular Southeast Asia, and Oceania, and is commonly associated with the Neolithic expansion of early Austronesian and Austroasiatic speakers that occurred more than 4,000 years ago.

References

  1. ↑ Huang, Xiufeng; Xia, Zi-Yang; Bin, Xiaoyun; He, Guanglin; Guo, Jianxin; Adnan, Atif; Yin, Lianfei; Huang, Youyi; Zhao, Jing; Yang, Yidong; Ma, Fuwei; Li, Yingxiang; Hu, Rong; Yang, Tianle; Wei, Lan-Hai; Wang, Chuan-Chao (30 June 2022). "Genomic Insights Into the Demographic History of the Southern Chinese". Frontiers in Ecology and Evolution. 10. doi: 10.3389/fevo.2022.853391 .
  2. ↑ Seguin-Orlando, Andaine; Korneliussen, Thorfinn S.; Sikora, Martin; Malaspinas, Anna-Sapfo; Manica, Andrea; Moltke, Ida; Albrechtsen, Anders; Ko, Amy; Margaryan, Ashot; Moiseyev, Vyacheslav; Goebel, Ted; Westaway, Michael; Lambert, David; Khartanovich, Valeri; Wall, Jeffrey D.; Nigst, Philip R.; Foley, Robert A.; Lahr, Marta Mirazon; Nielsen, Rasmus; Orlando, Ludovic; Willerslev, Eske (28 November 2014). "Genomic structure in Europeans dating back at least 36,200 years". Science. 346 (6213): 1113–1118. Bibcode:2014Sci...346.1113S. doi:10.1126/science.aaa0114. PMID   25378462. S2CID   206632421.
  3. ↑ Vallini, Leonardo; Zampieri, Carlo; Shoaee, Mohamed Javad; Bortolini, Eugenio; Marciani, Giulia; Aneli, Serena; Pievani, Telmo; Benazzi, Stefano; Barausse, Alberto; Mezzavilla, Massimo; Petraglia, Michael D.; Pagani, Luca (25 March 2024). "The Persian plateau served as hub for Homo sapiens after the main out of Africa dispersal". Nature Communications. 15 (1): 1882. Bibcode:2024NatCo..15.1882V. doi:10.1038/s41467-024-46161-7. PMC   10963722 . PMID   38528002.
  4. 1 2 Aoki, Kenichi; Takahata, Naoyuki; Oota, Hiroki; Wakano, Joe Yuichiro; Feldman, Marcus W. (30 August 2023). "Infectious diseases may have arrested the southward advance of microblades in Upper Palaeolithic East Asia". Proceedings of the Royal Society B: Biological Sciences. 290 (2005). doi:10.1098/rspb.2023.1262. PMC   10465978 . PMID   37644833. A single major migration of modern humans into the continents of Asia and Sahul was strongly supported by earlier studies using mitochondrial DNA, the non-recombining portion of Y chromosomes, and autosomal SNP data [42–45]. Ancestral Ancient South Indians with no West Eurasian relatedness, East Asians, Onge (Andamanese hunter–gatherers) and Papuans all derive in a short evolutionary time from the eastward dispersal of an out-of-Africa population [46,47]. [...] The HUGO (Human Genome Organization) Pan-Asian SNP consortium [44] investigated haplotype diversity within present-day Asian populations and found a strong correlation with latitude, with diversity decreasing from south to north. The correlation continues to hold when only mainland Southeast Asian and East Asian populations are considered, and is perhaps attributable to a serial founder effect [50]. These observations are consistent with the view that soon after the single eastward migration of modern humans, East Asians diverged in southern East Asia and dispersed northward across the continent.
  5. ↑ Moreno-Mayar, J. Víctor; Potter, Ben A.; Vinner, Lasse; Steinrücken, Matthias; Rasmussen, Simon; Terhorst, Jonathan; Kamm, John A.; Albrechtsen, Anders; Malaspinas, Anna-Sapfo; Sikora, Martin; Reuther, Joshua D.; Irish, Joel D.; Malhi, Ripan S.; Orlando, Ludovic; Song, Yun S.; Nielsen, Rasmus; Meltzer, David J.; Willerslev, Eske (2018), "Terminal Pleistocene Alaskan genome reveals first founding population of Native Americans" (PDF), Nature, 553 (7687): 203–207, Bibcode:2018Natur.553..203M, doi:10.1038/nature25173, PMID   29323294, S2CID   4454580
  6. ↑ Yang, Melinda A. (2022-01-06). "A genetic history of migration, diversification, and admixture in Asia". Human Population Genetics and Genomics. 2 (1): 1–32. doi: 10.47248/hpgg2202010001 . ISSN   2770-5005.
  7. ↑ Adhikari, Kaustubh; Fuentes-Guajardo, Macarena; Quinto-Sánchez, Mirsha; Mendoza-Revilla, Javier; Camilo Chacón-Duque, Juan; Acuña-Alonzo, Victor; Jaramillo, Claudia; Arias, William; Lozano, Rodrigo Barquera; Pérez, Gastón Macín; Gómez-Valdés, Jorge; Villamil-Ramírez, Hugo; Hunemeier, Tábita; Ramallo, Virginia; Silva de Cerqueira, Caio C. (2016-05-19). "A genome-wide association scan implicates DCHS2, RUNX2, GLI3, PAX1 and EDAR in human facial variation". Nature Communications. 7 (1): 11616. Bibcode:2016NatCo...711616A. doi:10.1038/ncomms11616. ISSN   2041-1723. PMC   4874031 . PMID   27193062.
  8. ↑ Yang, Melinda A. (2022-01-06). "A genetic history of migration, diversification, and admixture in Asia". Human Population Genetics and Genomics. 2 (1): 1–32. doi: 10.47248/hpgg2202010001 . ISSN   2770-5005. ...In contrast, mainland East and Southeast Asians and other Pacific islanders (e.g., Austronesian speakers) are closely related to each other [9,15,16] and here denoted as belonging to an East and Southeast Asian (ESEA) lineage (Box 2). …the ESEA lineage differentiated into at least three distinct ancestries: Tianyuan ancestry which can be found 40,000-33,000 years ago in northern East Asia, ancestry found today across present-day populations of East Asia, Southeast Asia, and Siberia, but whose origins are unknown, and Hòabìnhian ancestry found 8,000-4,000 years ago in Southeast Asia, but whose origins in the Upper Paleolithic are unknown.
  9. ↑ https://www.researchgate.net/publication/346962622_Fine-scale_north-to-south_genetic_admixture_profile_in_Shaanxi_Han_Chinese_revealed_by_genome-wide_demographic_history_reconstruction [ bare URL ]
  10. 1 2 3 He, Guang-Lin; Wang, Meng-Ge; Li, Ying-Xiang; et al. (2022). "Fine-scale north-to-south genetic admixture profile in Shaanxi Han Chinese revealed by genome-wide demographic history reconstruction". Journal of Systematics and Evolution. 60 (4): 955–972. doi:10.1111/jse.12715 – via Wiley Online Library.
  11. ↑ Chiang, Charleston W.K.; Mangul, Serghei; Robles, Christopher; Sankararaman, Sriram (2018). "A Comprehensive Map of Genetic Variation in the World's Largest Ethnic Group —Han Chinese". Molecular Biology and Evolution. 35 (11): 2736–2750. doi:10.1093/molbev/msy170. PMC   6693441 . PMID   30169787.
  12. ↑ Zhou, Jingbin; Zhang, Xianpeng; Li, Xin; et al. (2022). "Genetic structure and demographic history of Northern Han people in Liaoning Province inferred from genome-wide array data". Frontiers in Ecology and Evolution. 10. doi: 10.3389/fevo.2022.1014024 .
  13. ↑ Wang, Chuan-Chao; Yeh, Hui-Yuan; Popov, Alexander N.; Zhang, Hu-Qin; Matsumura, Hirofumi (2021-02-22). "Genomic insights into the formation of human populations in East Asia". Nature. 591 (7850). Springer: 413–419. Bibcode:2021Natur.591..413W. doi:10.1038/s41586-021-03336-2. ISSN   0028-0836. PMC   7993749 . PMID   33618348.
  14. ↑ Huang, Xiufeng; Xia, Zi-Yang; Bin, Xiaoyun; He, Guanglin; Guo, Jianxin (2022-06-30). "Genomic Insights Into the Demographic History of the Southern Chinese". Frontiers in Ecology and Evolution. 10. Frontiers Media SA. doi: 10.3389/fevo.2022.853391 . ISSN   2296-701X.
  15. ↑ Wang, Tianyi; Wang, Wei; Xie, Guangmao; Li, Zhen; Fan, Xuechun (2021). "Human population history at the crossroads of East and Southeast Asia since 11,000 years ago". Cell. 184 (14). Elsevier BV: 3829–3841.e21. doi: 10.1016/j.cell.2021.05.018 . ISSN   0092-8674. PMID   34171307.
  16. ↑ Soares, Pedro A.; Trejaut, Jean A.; Rito, Teresa; Cavadas, Bruno; Hill, Catherine; Eng, Ken Khong; Mormina, Maru; Brandão, Andreia; Fraser, Ross M.; Wang, Tse-Yi; Loo, Jun-Hun; Snell, Christopher; Ko, Tsang-Ming; Amorim, António; Pala, Maria; MacAulay, Vincent; Bulbeck, David; Wilson, James F.; Gusmão, Leonor; Pereira, Luísa; Oppenheimer, Stephen; Lin, Marie; Richards, Martin B. (2016). "Resolving the ancestry of Austronesian-speaking populations". Human Genetics. 135 (3): 309–326. doi: 10.1007/s00439-015-1620-z . PMC   4757630 . PMID   26781090.
  17. ↑ Hallast, Pille; Agdzhoyan, Anastasia; Balanovsky, Oleg; Xue, Yali; Tyler-Smith, Chris (2021-02-01). "A Southeast Asian origin for present-day non-African human Y chromosomes". Human Genetics. 140 (2): 299–307. doi:10.1007/s00439-020-02204-9. ISSN   1432-1203. PMC   7864842 . PMID   32666166.
  18. ↑ Bae, Christopher J.; Douka, Katerina; Petraglia, Michael D. (2017-12-08). "On the origin of modern humans: Asian perspectives". Science. 358 (6368): eaai9067. doi: 10.1126/science.aai9067 . ISSN   0036-8075. PMID   29217544. S2CID   4436271.
  19. ↑ "Asian Ancestry based on Studies of Y-DNA Variation: Part 1 Early origins — roots from Africa and emergence in East Asia". Genebase Tutorials. Archived from the original on 2012-01-02. Retrieved 2011-12-28.
  20. ↑ Shi H, Zhong H, Peng Y, et al. (2008). "Y chromosome evidence of earliest modern human settlement in East Asia and multiple origins of Tibetan and Japanese populations". BMC Biol. 6: 45. doi: 10.1186/1741-7007-6-45 . PMC   2605740 . PMID   18959782.
  21. ↑ WANG, Chuan-Chao; YAN, Shi; QIN, Zhen-Dong; LU, Yan; DING, Qi-Liang; WEI, Lan-Hai; LI, Shi-Lin; YANG, Ya-Jun; JIN, Li; LI, Hui (May 2013). "Late Neolithic expansion of ancient Chinese revealed by Y chromosome haplogroup O3a1c-002611". Journal of Systematics and Evolution. 51 (3): 280–286. doi: 10.1111/j.1759-6831.2012.00244.x . S2CID   55081530.
  22. ↑ https://docslide.net/documents/relocation-of-civilization-centers-in-ancient-china-environmental-factors.html%7C "Relocation of Civilization Centers in Ancient China: Environmental Factors"
  23. 1 2 Zhao, Yong-Bin; Zhang, Ye; Zhang, Quan-Chao; Li, Hong-Jie; Cui, Ying-Qiu; Xu, Zhi; Jin, Li; Zhou, Hui; Zhu, Hong (4 May 2015). "Ancient DNA Reveals That the Genetic Structure of the Northern Han Chinese Was Shaped Prior to 3,000 Years Ago". PLOS ONE. 10 (5): e0125676. Bibcode:2015PLoSO..1025676Z. doi: 10.1371/journal.pone.0125676 . PMC   4418768 . PMID   25938511.
  24. ↑ Gan RJ, Pan SL, Mustavich LF, Qin ZD, Cai XY, Qian J, Liu CW, Peng JH, Li SL, Xu JS, Jin L, Li H (2008). "Pinghua population as an exception of Han Chinese's coherent genetic structure". Journal of Human Genetics. 53 (4): 303–13. doi: 10.1007/s10038-008-0250-x . PMID   18270655.
  25. 1 2 3 Hammer MF, Karafet TM, Park H, Omoto K, Harihara S, Stoneking M, Horai S (2006). "Dual origins of the Japanese: common ground for hunter-gatherer and farmer Y chromosomes". Journal of Human Genetics. 51 (1): 47–58. doi: 10.1007/s10038-005-0322-0 . PMID   16328082.
  26. 1 2 3 4 5 Xue Y, Zerjal T, Bao W, Zhu S, Shu Q, Xu J, Du R, Fu S, Li P, Hurles ME, Yang H, Tyler-Smith C (April 2006). "Male demography in East Asia: a north-south contrast in human population expansion times". Genetics. 172 (4): 2431–9. doi:10.1534/genetics.105.054270. PMC   1456369 . PMID   16489223.
  27. 1 2 Kim SH, Kim KC, Shin DJ, Jin HJ, Kwak KD, Han MS, Song JM, Kim W, Kim W (April 2011). "High frequencies of Y-chromosome haplogroup O2b-SRY465 lineages in Korea: a genetic perspective on the peopling of Korea". Investigative Genetics. 2 (1): 10. doi: 10.1186/2041-2223-2-10 . PMC   3087676 . PMID   21463511. CC-BY icon.svg Text was copied from this source, which is available under a Creative Commons Attribution 2.0 (CC BY 2.0) license.
  28. 1 2 Yin C, Su K, He Z, Zhai D, Guo K, Chen X, Jin L, Li S (July 2020). "Genetic Reconstruction and Forensic Analysis of Chinese Shandong and Yunnan Han Populations by Co-Analyzing Y Chromosomal STRs and SNPs". Genes. 11 (7): 743. doi: 10.3390/genes11070743 . PMC   7397191 . PMID   32635262.
  29. ↑ Yao, H., Wang, M., Zou, X. et al., "New insights into the fine-scale history of western–eastern admixture of the northwestern Chinese population in the Hexi Corridor via genome-wide genetic legacy." Mol Genet Genomics 296, 631–651 (2021). https://doi.org/10.1007/s00438-021-01767-0
  30. ↑ Yao H, Wang M, Zou X, Li Y, Yang X, Li A, Yeh HY, Wang P, Wang Z, Bai J, Guo J, Chen J, Ding X, Zhang Y, Lin B, Wang CC, He G (May 2021). "New insights into the fine-scale history of western-eastern admixture of the northwestern Chinese population in the Hexi Corridor via genome-wide genetic legacy". Molecular Genetics and Genomics. 296 (3): 631–651. doi:10.1007/s00438-021-01767-0. PMID   33650010. S2CID   232091731.
  31. 1 2 3 Yan S, Wang CC, Li H, Li SL, Jin L (September 2011). "An updated tree of Y-chromosome Haplogroup O and revised phylogenetic positions of mutations P164 and PK4". European Journal of Human Genetics. 19 (9): 1013–5. doi:10.1038/ejhg.2011.64. PMC   3179364 . PMID   21505448.
  32. ↑ Katoh T, Munkhbat B, Tounai K, Mano S, Ando H, Oyungerel G, et al. (February 2005). "Genetic features of Mongolian ethnic groups revealed by Y-chromosomal analysis". Gene. 346: 63–70. doi:10.1016/j.gene.2004.10.023. PMID   15716011.
  33. 1 2 Su B, Xiao J, Underhill P, Deka R, Zhang W, Akey J, Huang W, Shen D, Lu D, Luo J, Chu J, Tan J, Shen P, Davis R, Cavalli-Sforza L, Chakraborty R, Xiong M, Du R, Oefner P, Chen Z, Jin L (December 1999). "Y-Chromosome evidence for a northward migration of modern humans into Eastern Asia during the last Ice Age". American Journal of Human Genetics. 65 (6): 1718–24. doi:10.1086/302680. PMC   1288383 . PMID   10577926.
  34. 1 2 Tajima A, Hayami M, Tokunaga K, Juji T, Matsuo M, Marzuki S, Omoto K, Horai S (2004). "Genetic origins of the Ainu inferred from combined DNA analyses of maternal and paternal lineages". Journal of Human Genetics. 49 (4): 187–93. doi: 10.1007/s10038-004-0131-x . PMID   14997363.
  35. 1 2 3 4 Trejaut JA, Poloni ES, Yen JC, Lai YH, Loo JH, Lee CL, He CL, Lin M (June 2014). "Taiwan Y-chromosomal DNA variation and its relationship with Island Southeast Asia". BMC Genetics. 15: 77. doi: 10.1186/1471-2156-15-77 . PMC   4083334 . PMID   24965575.
  36. ↑ Yan S, Wang CC, Zheng HX, Wang W, Qin ZD, Wei LH, et al. (2014). "Y chromosomes of 40% Chinese descend from three Neolithic super-grandfathers". PLOS ONE. 9 (8): e105691. arXiv: 1310.3897 . Bibcode:2014PLoSO...9j5691Y. doi: 10.1371/journal.pone.0105691 . PMC   4149484 . PMID   25170956.
  37. ↑ Hurles ME, Sykes BC, Jobling MA, Forster P (May 2005). "The dual origin of the Malagasy in Island Southeast Asia and East Africa: evidence from maternal and paternal lineages". American Journal of Human Genetics. 76 (5): 894–901. doi:10.1086/430051. PMC   1199379 . PMID   15793703.
  38. ↑ Sun, Na; Ma, Peng-Cheng; Yan, Shi; et al. (2019). "Phylogeography of Y-chromosome haplogroup Q1a1a-M120, a paternal lineage connecting populations in Siberia and East Asia". Annals of Human Biology. 46 (3): 261–266. doi:10.1080/03014460.2019.1632930. PMID   31208219 – via Taylor & Francis Online.
  39. ↑ Wang, Yuchen; Lu, Dongsheng; Chung, Yeun-Jun; Xu, Shuhua (2018-04-06). "Genetic structure, divergence and admixture of Han Chinese, Japanese and Korean populations". Hereditas. 155 (1): 19. doi: 10.1186/s41065-018-0057-5 . ISSN   1601-5223. PMC   5889524 . PMID   29636655.
  40. ↑ Yanan Cao; Lin Li; Min Xu; Zhimin Feng; Xiaohui Sun; Jieli Lu; Yu Xu; Peina Du; Tiange Wang; Ruying Hu; Zhen Ye; Lixin Shi; Xulei Tang; Li Yan (30 April 2020). "The ChinaMAP analytics of deep whole genome sequences in 10,588 individuals". Cell Research. 30 (9): 717–731. doi:10.1038/s41422-020-0322-9. PMC   7609296 . PMID   32355288.
  41. ↑ Wang, Yuchen; Lu, Dongsheng; Chung, Yeun-Jun; Xu, Shuhua (6 April 2018). "Genetic structure, divergence and admixture of Han Chinese, Japanese and Korean populations". Hereditas. 155 (19): 19. doi: 10.1186/s41065-018-0057-5 . PMC   5889524 . PMID   29636655.
  42. ↑ Wen B, Li H, Lu D, Song X, Zhang F, He Y, et al. (September 2004). "Genetic evidence supports demic diffusion of Han culture". Nature. 431 (7006): 302–305. Bibcode:2004Natur.431..302W. doi:10.1038/nature02878. PMID   15372031. S2CID   4301581.
  43. ↑ Zhao YB, Zhang Y, Zhang QC, Li HJ, Cui YQ, Xu Z, Jin L, Zhou H, Zhu H (2015). "Ancient DNA reveals that the genetic structure of the northern Han Chinese was shaped prior to 3,000 years ago". PLOS ONE. 10 (5): e0125676. Bibcode:2015PLoSO..1025676Z. doi: 10.1371/journal.pone.0125676 . PMC   4418768 . PMID   25938511.
  44. ↑ Yao YG, Kong QP, Bandelt HJ, Kivisild T, Zhang YP (March 2002). "Phylogeographic differentiation of mitochondrial DNA in Han Chinese". American Journal of Human Genetics. 70 (3): 635–51. doi:10.1086/338999. PMC   384943 . PMID   11836649.
  45. ↑ Kivisild T, Tolk HV, Parik J, Wang Y, Papiha SS, Bandelt HJ, Villems R (October 2002). "The emerging limbs and twigs of the East Asian mtDNA tree". Molecular Biology and Evolution. 19 (10): 1737–51. doi: 10.1093/oxfordjournals.molbev.a003996 . PMID   12270900.
  46. ↑ Yao YG, Kong QP, Man XY, Bandelt HJ, Zhang YP (February 2003). "Reconstructing the evolutionary history of China: a caveat about inferences drawn from ancient DNA". Molecular Biology and Evolution. 20 (2): 214–9. doi: 10.1093/molbev/msg026 . PMID   12598688.
  47. ↑ Kong QP, Sun C, Wang HW, Zhao M, Wang WZ, Zhong L, Hao XD, Pan H, Wang SY, Cheng YT, Zhu CL, Wu SF, Liu LN, Jin JQ, Yao YG, Zhang YP (January 2011). "Large-scale mtDNA screening reveals a surprising matrilineal complexity in east Asia and its implications to the peopling of the region". Molecular Biology and Evolution. 28 (1): 513–22. doi: 10.1093/molbev/msq219 . PMID   20713468.
  48. ↑ Zhang Chi; Hsiao-Chun Hung (Fall 2008). "The Neolithic of Southern China-Origin, Development, and Dispersal". Asian Perspectives. 47 (2): 299–329. doi:10.1353/asi.0.0004. hdl: 10125/17291 . S2CID   162258899.
  49. ↑ Shelach, Gideon (December 2000). "The Earliest Neolithic Cultures of Northeast China: Recent Discoveries and New Perspectives on the Beginning of Agriculture". Journal of World Prehistory. 14 (4): 363–413. doi:10.1023/A:1011124209079. S2CID   161599532.
  50. ↑ Zhang Chi; Hsiao-chun Hun (March 2010). "The emergence of agriculture in southern China". Antiquity. 84 (323): 11–25. doi:10.1017/S0003598X00099737. S2CID   87789369.
  51. ↑ Higham, C.F.W.; Xie Guangmao; Lin Qiang (June 2011). "The prehistory of a Friction Zone: first farmers and hunters-gatherers in Southeast Asia". Antiquity. 85 (328): 529–543. doi:10.1017/S0003598X00067922. S2CID   162768159.
  52. ↑ Aldenderfer, Mark; Zhang Yinong (March 2004). "The Prehistory of the Tibetan Plateau to the Seventh Century A.D.: Perspectives and Research from China and the West Since 1950". Journal of World Prehistory. 18 (1): 1–55. doi:10.1023/B:JOWO.0000038657.79035.9e. S2CID   154022638.
  53. ↑ Blench, Roger. 2009. If agriculture cannot be reconstructed for Proto-Sino-Tibetan, what are the consequences? Archived 2019-05-05 at the Wayback Machine . Paper presented at the 42nd International Conference on Sino-Tibetan Language and Linguistics (ICSTLL 42), Chiang Mai, November 2–4, 2009. (slides)
  54. ↑ Sagart, Laurent; Jacques, Guillaume; Lai, Yunfan; Ryder, Robin; Thouzeau, Valentin; Greenhill, Simon J.; List, Johann-Mattis (2019). "Dated language phylogenies shed light on the ancestry of Sino-Tibetan". Proceedings of the National Academy of Sciences of the United States of America. 116 (21): 10317–10322. Bibcode:2019PNAS..11610317S. doi: 10.1073/pnas.1817972116 . PMC   6534992 . PMID   31061123.
  55. ↑ Black ML, Wise CA, Wang W, Bittles AH (June 2006). "Combining genetics and population history in the study of ethnic diversity in the People's Republic of China". Hum. Biol. 78 (3): 277–93. doi:10.1353/hub.2006.0041. PMID   17216801. S2CID   42002729.
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