| Clark's anemonefish | |
|---|---|
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| Clark's anemonefish with sea anemone | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| Family: | Pomacentridae |
| Genus: | Amphiprion |
| Species: | A. clarkii |
| Binomial name | |
| Amphiprion clarkii (J. W. Bennett, 1830) | |
| Synonyms | |
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Clark's anemonefish (Amphiprion clarkii), also known as the yellowtail clownfish, is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes. [3]
Clownfish or anemonefish are fishes that, in the wild, form symbiotic mutualisms with sea anemones and are unaffected by the stinging tentacles of the host anemone, see Amphiprioninae § Symbiosis and mutualism. The sea anemone protects the clownfish from predators, as well as providing food through the scraps left from the anemone's meals and occasional dead anemone tentacles. In return, the clownfish defends the anemone from its predators, and parasites. [4] Clownfish are small-sized, 10–18 centimetres (3.9–7.1 in), and depending on species, they are overall yellow, orange, or a reddish or blackish color, and many show white bars or patches. Within species there may be color variations, most commonly according to distribution, but also based on sex, age and host anemone. Clownfish are found in warmer waters of the Indian and Pacific oceans and the Red Sea in sheltered reefs or in shallow lagoons.
In a group of clownfish, there is a strict dominance hierarchy. The largest and most aggressive fish is female and is found at the top. Only two clownfish, a male and a female, in a group reproduce through external fertilization. Clownfish are sequential hermaphrodites, meaning that they develop into males first, and when they mature, they become females. They are not aggressive. When the egg production and spawning patterns of eight breeding pairs of yellowtail clownfish, Amphiprion clarkii, were observed in a coral reef off the coast of the Philippines, they preferred breeding in the colder months. A trend of a peak breeding season from November through May was seen; egg production increased up to the new moon and decreased after the full moon. This seasonality of spawning is comparable to other species of clownfish that live in temperate regions (Holtswarth & San Jose, 2017).
Clark's anemonefish is a small-sized fish which grows up to 10 cm as a male and 15 cm as a female. [5] [6] It is stocky, laterally compressed, and oval to rounded.
It is colorful, with vivid black, white, and yellow stripes, though the exact pattern shows considerable geographical variation. Usually it is black dorsally and orange-yellow ventrally, the black areas becoming wider with age. [7] There are two vertical white bands, one behind the eye and one above the anus, and the caudal peduncle is white. The snout is orange or pinkish. The dorsal and caudal fins are orange-yellow, [8] and the caudal fin is generally lighter in tone than the rest of the body, sometimes becoming whitish. [9]
Clark's anemonefish shows the greatest color variations of any anemonefish, with variations based on location, sex, age and host anemone. Adults in Vanuatu and New Caledonia are orange-yellow with two vertical white bands. [10] Sex related color differences may be present, such as the female having a white caudal fin and the male having a yellow caudal fin. [10] Juveniles are orange-yellow with vertical white bands. [8] Fish living with the host anemone Stichodactyla mertensii , Mertens' carpet sea anemone are frequently black except for the snout bars and tail. [10]
The caudal fin is forked and the base lacks a white bar on A. latifasciatus . The caudal fin lacks the sharp demarcation between white and dark and the mid-body bar is narrower on A. allardi and A. akindynos . The caudal fin is dark on A. chrysogaster , A. fuscocaudatus and A. tricinctus . [10]
Clark's anemonefish is the most widely distributed anemonefish, being found in tropical waters from the Indian Ocean to the Western Pacific. [9]
Clark's anemonefish is the least host specific anemonefish, living in association with all ten species of sea anemones that host anemonefish:
The specific name and the common name both honour the Scottish engraver John Heaviside Clark (1771–1836) who provided illustrations for Bennett's A Selection from the most remarkable and interesting Fishes found on the Coast of Ceylon, from drawings made in the Southern part of that Island. [11]
Heteractis magnifica, also known by the common names magnificent sea anemone or Ritteri anemone, is a species of sea anemone belonging to the Stichodactylidae family native to the Indo-Pacific area.
The ocellaris clownfish, also known as the false percula clownfish or common clownfish, is a marine fish belonging to the family Pomacentridae, which includes clownfishes and damselfishes. Amphiprion ocellaris are found in different colors, depending on where they are located. For example, black Amphiprion ocellaris with white bands can be found near northern Australia, Southeast Asia, and Japan. Orange or red-brown Amphiprion ocellaris also exist with three similar white bands on the body and head. Amphiprion ocellaris can be distinguished from other Amphiprion species based on the number of pectoral rays and dorsal spines. Amphiprion ocellaris are known to grow about 110 mm long. Like many other fish species, females are, however, larger than males. The life cycle of Amphiprion ocellaris varies in whether they reside at the surface or bottom of the ocean. When they initially hatch, they reside near the surface. However, when Amphiprion ocellaris enter into the juvenile stage of life, they travel down to the bottom to find shelter in a host anemone. Once they find their anemone, they form a symbiotic relationship with them.
Amphiprion sandaracinos, also known as the orange skunk clownfish or orange anemonefish, is a species of anemonefish that is distinguished by its broad white stripe along the dorsal ridge. Like all anemonefishes it forms a symbiotic mutualism with sea anemones and is unaffected by the stinging tentacles of the host anemone. It is a sequential hermaphrodite with a strict sized based dominance hierarchy: the female is largest, the breeding male is second largest, and the male non-breeders get progressively smaller as the hierarchy descends. They exhibit protandry, meaning the breeding male will change to female if the sole breeding female dies, with the largest non-breeder becomes the breeding male.
Amphiprion sebae, also known as the sebae clownfish, is an anemonefish found in the northern Indian Ocean, from Java to the Arabian Peninsula. Like all anemonefish it is usually found living in association with sea anemones. While the common name of Heteractis crispa, the sebae anemone, suggests an association, it is normally found with the Stichodactyla haddoni or saddle anemone. A. sebae, like all anemonefish, lives in a symbiotic relationship with the host anemone where the fish is unaffected by the stinging tentacles of the anemone. In a group of clownfish, Only two clownfish, a male and a female, in a group reproduce through external fertilization. Clownfish are sequential hermaphrodites, changing from male to female, with a strict dominance hierarchy and only the largest fish being female.
The orange-fin anemonefish is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes, found in the Western Pacific north of the Great Barrier Reef from the surface to 20 m, to include the Pacific Ocean between Queensland, Australia, and New Guinea to the Marshall and Tuamotus Islands. It can grow to 17 cm in length.
Allard's clownfish or Allard's anemonefish is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes, from the western Indian Ocean off the coast of East Africa and the Mascarenes.
Amphiprion polymnus, also known as the saddleback clownfish or yellowfin anemonefish, is a black and white species of anemonefish with a distinctive saddle. Like all anemonefishes it forms a symbiotic mutualism with sea anemones and is unaffected by the stinging tentacles of the host anemone. It is a sequential hermaphrodite with a strict sized-based dominance hierarchy: the female is largest, the breeding male is second largest, and the male non-breeders get progressively smaller as the hierarchy descends. They exhibit protandry, meaning the breeding male will change to female if the sole breeding female dies, with the largest non-breeder becomes the breeding male.
Amphiprion akindynos, the Barrier Reef anemonefish, is a species of anemonefish that is principally found in the Great Barrier Reef of Australia, but also in nearby locations in the Western Pacific. The species name 'akindynos' is Greek, meaning 'safe' or 'without danger' in reference to the safety afforded amongst the tentacles of its host anemone. Like all anemonefishes it forms a symbiotic mutualism with sea anemones and is unaffected by the stinging tentacles of the host anemone. It is a sequential hermaphrodite with a strict size-based dominance hierarchy: the male is are largest, the breeding male is second largest, and the male non-breeders get progressively smaller as the hierarchy descends. They exhibit protandry, meaning the breeding male will change to female if the sole breeding female dies, with the largest non-breeder becomes the breeding male. The fish's natural diet includes zooplankton.
The Red Sea Clownfish, commonly known as the Red Sea or two-bandedanemonefish is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes. Like other species of the genus, the fish feeds on algae and zooplankton in the wild.
The pink skunk clownfish, also known as the pink anemonefish, is a species of anemonefish that is widespread from northern Australia through the Malay Archipelago and Melanesia. Like all anemonefishes, it forms a symbiotic mutualism with sea anemones and is unaffected by the stinging tentacles of the host. It is a sequential hermaphrodite with a strict size-based dominance hierarchy; the female is largest, the breeding male is second largest, and the male nonbreeders get progressively smaller as the hierarchy descends. They exhibit protandry, meaning the breeding male changes to female if the sole breeding female dies, with the largest nonbreeder becoming the breeding male.
The three-band anemonefish is a species of anemonefish endemic to the Marshall Islands in the western part of the Pacific Ocean. Like all anemonefishes, it forms a symbiotic mutualism with sea anemones and is unaffected by the stinging tentacles of its host. It is a sequential hermaphrodite with a strict size-based dominance hierarchy; the female is largest, the breeding male is second largest, and the male nonbreeders get progressively smaller as the hierarchy descends. They exhibit protandry, meaning the breeding male changes to female if the sole breeding female dies, with the largest nonbreeder becoming the breeding male. The fish's natural diet includes zooplankton.
The nosestripe clownfish or nosestripe anemonefish, skunk clownfish, Amphiprion akallopisos, is an anemonefish that lives in association with sea anemones. A. akallopisos is found in the Indian Ocean. It resides in shallow inshore reefs as deep as 15 m with a moderate to strong current. The skunk clownfish can also be kept in captivity by aquarists.
Stichodactyla mertensii, commonly known as Mertens' carpet sea anemone, is a species of sea anemones in the family Stichodactylidae. It is regarded as the largest sea anemone with a diameter of over 1 m (3.3 ft), the next largest being Heteractis magnifica, which has longer tentacles. This species has an oral disc that can be described as more ovoid than circular that contours to the surrounding substrate and is attached to the substrate by adhesive verrucae, which are wart-like projections. Its blunt or pointed tentacles are uniformly shaped, and are only about 1–2 centimetres (0.39–0.79 in) long. It contains obligate symbiotic zooxanthellae, and is a host to around half the species of anemonefish and one damselfish, Dascyllus trimaculatus.
Amphiprion chagosensis, the Chagos anemonefish, is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes. It is named for the Chagos Archipelago in the Indian Ocean and it is endemic to the archipelago. The original specimens were collected at Diego Garcia Atoll, Chagos Archipelago.
Amphiprion chrysogaster, the Mauritian anemonefish, is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes. It is endemic to Mauritius and probably Réunion.
Amphiprion fuscocaudatus is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes.
Amphiprion latifasciatus is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes.
Amphiprion leucokranos is a naturally occurring hybrid anemonefish found in the western central Pacific Ocean. Like all anemonefishes it forms a symbiotic mutualism with sea anemones and is unaffected by the stinging tentacles of the host anemone. It is a sequential hermaphrodite with a strict dominance hierarchy, features which are critical to the direction of gene flow.
Amphiprion omanensis is a marine fish belonging to the family Pomacentridae, the clownfishes and damselfishes.
Amphiprion thiellei is thought to be a naturally occurring hybrid anemonefish found in the vicinity of Cebu, Philippines. Like all anemonefishes it forms a symbiotic mutualism with sea anemones and is unaffected by the stinging tentacles of the host anemone. It is a sequential hermaphrodite with a strict dominance hierarchy.
What size? Males maximum 4 in (10 cm), females maximum 5 7/8 in (15 cm)
Media related to Amphiprion clarkii at Wikimedia Commons
Data related to Amphiprion clarkii at Wikispecies
