The billfish are a group of saltwater predatory fish characterised by prominent pointed bills (rostra), and by their large size; some are longer than 4 m (13 ft). Extant billfish include sailfish and marlin, which make up the family Istiophoridae; and swordfish, sole member of the family Xiphiidae. They are often apex predators which feed on a wide variety of smaller fish, crustaceans and cephalopods. These two families are sometimes classified as belonging to the order Istiophoriformes, a group which originated around 71 million years ago in the Late Cretaceous, with the two families diverging around 15 million years ago in the Late Miocene. However, they are also classified as being closely related to the mackerels and tuna within the suborder Scombroidei of the order Perciformes. However, the 5th edition of the Fishes of the World does recognise the Istiophoriformes as a valid order, albeit including the Sphyraenidae, the barracudas.
Protosphyraena is a fossil genus of swordfish-like marine fish, that thrived worldwide during the Upper Cretaceous Period (Coniacian-Maastrichtian). Though fossil remains of this taxon have been found in both Europe and Asia, it is perhaps best known from the Smoky Hill Member of the Niobrara Chalk Formation of Kansas. Protosphyraena was a large fish, averaging 2–3 metres in length. Protosphyraena shared the Cretaceous oceans with aquatic reptiles, such as mosasaurs and plesiosaurs, as well as with many other species of extinct predatory fish. The name Protosphyraena is a combination of the Greek word protos ("early") plus Sphyraena, the genus name for barracuda, as paleontologists initially mistook Protosphyraena for an ancestral barracuda. Recent research shows that the genus Protosphyraena is not at all related to the true swordfish-family Xiphiidae, but belongs to the extinct family Pachycormidae.
Anatalavis is genus of prehistoric birds related to ducks and geese, perhaps to the magpie-goose in particular. Alternatively, it might have been a more basal lineage of Anserimorphae distinct from the living waterfowl, similar or even related to the roughly contemporary Conflicto antarcticus from the Danian of Antarctica.
Bothremydidae is an extinct family of side-necked turtles (Pleurodira) known from the Cretaceous and Cenozoic. They are closely related to Podocnemididae, and are amongst the most widely distributed pleurodire groups, with their fossils having been found in Africa, India, the Middle East, Europe, North America and South America. Bothremydids were aquatic turtles with a high morphological diversity, indicative of generalist, molluscivorous, piscivorous and possibly herbivorous grazing diets, with some probably capable of suction feeding. Unlike modern pleurodires, which are exclusively freshwater, bothremydids inhabited freshwater, marine and coastal environments. Their marine habits allowed bothremydids to disperse across oceanic barriers into Europe and North America during the early Late Cretaceous (Cenomanian). The youngest records of the group are indeterminate remains from Saudi Arabia and Oman, dating to the Miocene.
Cimolichthys is an extinct genus of large predatory marine aulopiform fish known worldwide from the Late Cretaceous. It is the only member of the family Cimolichthyidae.
Blochius is an extinct genus of prehistoric billfish from the Eocene. It is only known from the Monte Bolca deposits in Italy, and was likely restricted to shallow, tropical waters of the Tethys Sea.
Stratodus is a genus of giant prehistoric aulopiform fish found in Cretaceous-aged marine strata of Kansas, Alabama, Morocco, Israel, and Niger, South Dakota, Jordan. It has also been found in the Tamaguélelt Formation of Mali, dating to the Lower Eocene, indicating that Stratodus survived the Cretaceous-Paleogene extinction event. This sleek fish has an upper jaw filled with multiple rows of tiny teeth and was the largest aulopiform, reaching 5 meters in length.
Acestrus is an extinct genus of prehistoric marine ray-finned fish that lived during the lower Eocene in Europe. It contains one species, A. ornatus from the London Clay, known from a single braincase. It is thought to possibly be closely allied with billfish based on the braincase morphology, although it remains uncertain whether it had the rostrum characteristic of billfishes. Some authorities have suggested blochiid affinities.
Aulopopsis is an extinct genus of prehistoric marine ray-finned fish that lived during the lower Eocene. It is considered a relative of lizardfish in the order Aulopiformes, but its exact taxonomic placement is uncertain. Some authorities place it with the Aulopidae, while others place it with the Giganturoidae.
Aglyptorhynchus is an extinct genus of prehistoric marine billfish that was distributed worldwide from the early Eocene to the early Miocene. Fossils are primarily known from the Northern Hemisphere, but one species is also known to have inhabited the waters off New Zealand.
Ampheristus is an extinct genus of prehistoric marine ray-finned fish. It was a basal or stem member of the family Ophidiidae, which contains modern cusk-eels. Fossils are known from worldwide from the Late Cretaceous to the late Paleogene, making it a rather successful survivor of the Cretaceous-Paleogene extinction event.
Brychaetus is an extinct genus of prehistoric marine bonytongue fish known from the Late Cretaceous to the late Eocene of Europe, North America, and northern Africa.
Coelodus is an extinct genus of marine and possibly freshwater pycnodont fish. It contains only one definitive species, C. saturnusHeckel, 1854, from the Late Cretaceous of Slovenia. Other species from the Late Jurassic to the Eocene have also been attributed to this genus based on isolated dental elements, but their assignment to Coelodus is uncertain, and this genus likely represents a non-monophyletic wastebasket taxon. A potential diagnostic trait is a prearticular tooth row with three regular highly elongated teeth.
Cylindracanthus is an extinct genus of Cretaceous to Eocene aged ray-finned fish. It is almost exclusively known from bony rostrum fragments as well as some associated teeth, with the rest of the skeleton being cartilaginous. While originally considered to be closely related to billfish, the structure of its rostrum is dissimilar, and is unlikely to be closely related, some later studies suggested closer affinities to the Acipenseriformes. However, this was later rejected due to the lack of osteocytes in histologically examined specimens, which resembles the condition of derived teleosts. A close relation to Blochius has been suggested.
Martinichthys is an extinct genus of plethodid fish from the Cretaceous of North America. It is known from the Niobrara Chalk, in which it is exceedingly rare. It is named after one H. T. Martin, who collected the most complete specimen at the time of description.
Cretalamna is a genus of extinct otodontid shark that lived from the latest Early Cretaceous to Eocene epoch. It is considered by many to be the ancestor of the largest sharks to have ever lived, such as Otodus angustidens, Otodus chubutensis, and Otodus megalodon.
Plethodidae is an extinct family of teleost fish that existed during the Late Cretaceous period. Fossils are known from North America, North Africa, and Europe.
Propristis is an extinct genus of sawfish that lived from the Eocene to the Miocene. It contains two valid species, P. schweinfurthi and P. mayumbensis. It has been found in Egypt, Cabinda, Morocco, Qatar, Spain, the United Kingdom, and the United States. Isolated rostral denticles are the most common remains, but rostra have also been found.
Rhamphoichthys is an extinct genus of billfish-like plethodid ray-finned fish from Late Cretaceous (Cenomanian). It contains one valid species, R. taxidiotis from the Hesseltal Formation of Germany and the Sannine Formation of Lebanon, in addition to two dubious species that are considered nomina nuda due to their fragmentary nature.
