Cretodus Temporal range: | |
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C. crassidens teeth from the Turonian of Italy | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Chondrichthyes |
Superorder: | Selachimorpha |
Order: | Lamniformes |
Family: | † Pseudoscapanorhynchidae |
Genus: | † Cretodus Sokolov, 1965 |
Type species | |
†Cretodus crassidens Dixon, 1850 | |
species | |
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Cretodus is an extinct genus of large mackerel sharks belonging to the proposed family Pseudoscapanorhynchidae. [1] Cretodus lived during the Late Cretaceous, ranging from the Cenomanian [2] to the Coniacian [3] (approximately 100 to 89 million years ago). The genus is well-known from strata deposited in the Western Interior Seaway (North America), [1] [2] [3] and from the Late Cretaceous of Europe, [4] Africa, and possibly Asia.[ citation needed ]Cretodus is primarily represented in the fossil record by isolated teeth and vertebral centra, though a couple of associated dentitions and vertebral columns have been found. [1] [4]
Prior to 2017, Cretodus was mainly known from isolated teeth. [5] The discovery of a virtually complete skeleton suggests the genus possessed a body similar in form to the modern tiger shark, with a wide head that was equipped with powerful jaws, and a stout body. [4] The morphology of Cretodus' placoid scales and vertebral centra suggest a moderate swimming speed, less than that of the fast-cruising contemporaneous macropredatory shark, Cretoxyrhina . [4] [1]
Size estimation is only reasonable for the few associated partial skeletons known. Shimada & Everhart (2019) reported an approximate total length of 5.15 m (16.9 ft) for the associated specimen of C. houghtonorum from Kansas which probably died at 22 years of age; they also suggested that the maximum total length of C. houghtonorum would have been 6.84 m (22.4 ft). [1] Amalfitano et al. (2022) reported an estimated total length of 6.6–7.8 m (22–26 ft) for a nearly complete specimen of C. crassidens from Italy which probably died at 23 years of age; they also used a growth model to estimate the maximum total length for C. crassidens at 9.55–11.28 m (31.3–37.0 ft). [4]
Cretodus teeth have lateral cusplets that are much smaller than the main cusp and well-separated from it, but connected by enamel on the labial face. The enamel surface is smooth, except near the base of the crown, where it forms vertical striae or costulae (grooves and ridges), a feature more prominent on the lingual face. The root is bilobate and has a massive shelf-like lingual root protuberance which diminishes in size in posterior positions. [4] [6]
It is estimated that C. houghtonorum was about 1.18 m (3.9 ft) long at birth, became sexually mature at 10 to 15 years of age, and reached its maximum lifespan at 51 to 55 years of age. [1] [4] C. crassidens was about 1.42 m (4.7 ft) long at birth, became sexually mature at 12 to 17 years of age, and reached its maximum life span at 64 years of age. [4]
Cretodus predated on large prey, and is suggested to have specialized in marine turtles, based on inferred stomach contents (a 2 m (6.6 ft) chelonioid turtle) of a specimen of C. crassidens found in Italy. [5] A predator-prey relationship has also been inferred between C. houghtonorum and hybodont sharks based on stomach contents. [1]
The distribution of fossils of C. crassidens suggests a largely cosmopolitan distribution and a preference for offshore areas in contrast with other species in the genus (e.g. C. houghtonorum), which preferred shallow water. [4]
Within the Western Interior Seaway, Cretodus lived among a number of other genera of shark including Cretoxyrhina, Squalicorax , Ptychodus, Cretalamna, Scapanorhynchus, and hybodont sharks. [1] In order to coexist, these sharks likely practiced resource partitioning, by occupying different regions (e.g. nearshore v.s. offshore) or specializing in different prey items. [4] [1]
Shimada & Everhart (2019) attributed the following species to the genus, asserting they were chronospecies belonging to three grades of crown broadness. [1]
Megalodon, meaning "big tooth", is an extinct species of mackerel shark that lived approximately 23 to 3.6 million years ago (Mya), from the Early Miocene to the Pliocene epochs. It was formerly thought to be a member of the family Lamnidae and a close relative of the great white shark. However, it is now classified into the extinct family Otodontidae, which diverged from the great white shark during the Early Cretaceous.
Squalicorax, commonly known as the crow shark, is a genus of extinct lamniform shark known to have lived during the Cretaceous period. The genus had a global distribution in the Late Cretaceous epoch. Multiple species within this genus are considered to be wastebasket taxon due to morphological similarities in the teeth.
Cretoxyrhina is an extinct genus of large mackerel shark that lived about 107 to 73 million years ago during the late Albian to late Campanian of the Late Cretaceous. The type species, C. mantelli, is more commonly referred to as the Ginsu shark, first popularized in reference to the Ginsu knife, as its theoretical feeding mechanism is often compared with the "slicing and dicing" when one uses the knife. Cretoxyrhina is traditionally classified as the likely sole member of the family Cretoxyrhinidae but other taxonomic placements have been proposed, such as within the Alopiidae and Lamnidae.
Xiphactinus is an extinct genus of large predatory marine bony fish that lived during the Late Cretaceous. Species in the genus bore a superficial resemblance to a gargantuan, fanged tarpon.
Tylosaurus is a genus of mosasaur, a large, predatory marine reptile closely related to modern monitor lizards and to snakes, from the Late Cretaceous.
Protostegidae is a family of extinct marine turtles that lived during the Cretaceous period. The family includes some of the largest sea turtles that ever existed. The largest, Archelon, had a head one metre (39 in) long. Like most sea turtles, they had flattened bodies and flippers for front appendages; protostegids had minimal shells like leatherback turtles of modern times.
Styxosaurus is a genus of plesiosaur of the family Elasmosauridae. Styxosaurus lived during the Campanian age of the Cretaceous period. Two species are known: S. snowii and S. browni.
Ptychodus is a genus of extinct durophagous (shell-crushing) sharks from the Late Cretaceous. Fossils of Ptychodus teeth are found in many Late Cretaceous marine sediments. There are many species among the Ptychodus that have been uncovered on all the continents around the globe. Such species are Ptychodus mortoni, P. decurrens, P. marginalis, P. mammillaris, P. rugosus and P. latissimus to name a few. They died out approximately 85 million years ago. A large number of remains have been found in the former Western Interior Seaway. A 2016 publication found that Ptychodus are likely true sharks belonging to Selachimorpha, rather than hybodonts or batoids as previously thought. Their life history coincides with the typical life of many other large sharks: they lived relatively long lives and were slow growing and produced large offspring and small litters.
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like chondrichthyans, which existed from the late Devonian to the Late Cretaceous. They form the group of Elasmobranchii closest to neoselachians, the clade of modern sharks and rays. Hybodonts were named and are distinguished based on their conical tooth shape. They are also noted for the presence of a spine on each of their two dorsal fins. They were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct.
During the time of the deposition of the Niobrara Chalk, much life inhabited the seas of the Western Interior Seaway. By this time in the Late Cretaceous many new lifeforms appeared such as mosasaurs, which were to be some of the last of the aquatic lifeforms to evolve before the end of the Mesozoic. Life of the Niobrara Chalk is comparable to that of the Dakota Formation, although the Dakota Formation, which was deposited during the Cenomanian, predates the chalk by about 10 million years.
The Carlile Shale is a Turonian age Upper/Late Cretaceous series shale geologic formation in the central-western United States, including in the Great Plains region of Colorado, Kansas, Nebraska, New Mexico, North Dakota, South Dakota, and Wyoming.
Otodus chubutensis, meaning "ear-shaped tooth of Chubut", from Ancient Greek ὠτ and ὀδούς – thus, "ear-shaped tooth", is an extinct species of prehistoric megatoothed sharks in the genus Otodus, that lived during Oligocene, Miocene, and Pliocene epochs, approximately 28 – 5 million years ago. The largest individuals were about 13.5 metres (44 ft) long. This shark is considered to be a close relative of the famous prehistoric megatoothed shark, O. megalodon. However, as is the case with O. megalodon, the classification of this species is disputed.
Cardabiodon is an extinct genus of large mackerel shark that lived about 95 to 91 million years ago (Ma) during the Cenomanian to Turonian of the Late Cretaceous. It is a member of the Cardabiodontidae, a family unique among mackerel sharks due to differing dental structures, and contains the two species C. ricki and C. venator. Cardabiodon fossils have been found in Australia, North America, England, and Kazakhstan. It was likely an antitropical shark that inhabited temperate neritic and offshore oceans between 40° and 60° paleolatitude, similar to the modern porbeagle shark.
Meristodonoides is an extinct genus of cartilaginous fish. The type species is M. rajkovichi, which was originally a species in the genus Hybodus. The species, along with other Hybodus species such as H. butleri and H. montanensis, was reassigned to Meristodonoides by Charlie J. Underwood and Stephen L. Cumbaa in 2010.
Cretalamna is a genus of extinct otodontid shark that lived from the latest Early Cretaceous to Eocene epoch. It is considered by many to be the ancestor of the largest sharks to have ever lived, Otodus angustidens, Otodus chubutensis, and Otodus megalodon.
Coniasaurus is an extinct genus of Late Cretaceous marine squamates that range in age from Cenomanian to Santonian. It was first described by Richard Owen in 1850 from lower Cenomanian chalk deposits in southeast England (Sussex). Two species have been described from this genus: C. crassidens, known from Cenomanian to Santonian deposits from southeast England, Germany and North America, and C. gracilodens from the Cenomanian of southeast England.
Mauriciosaurus is a genus of polycotylid plesiosaur from the Late Cretaceous of Mexico. It contains a single species, M. fernandezi, described in 2017 by Eberhard Frey and colleagues from a single well-preserved juvenile specimen about 1.9 metres long. Morphologically, it is overall most similar to the polycotyline polycotylids Trinacromerum and Dolichorhynchops. However, several features separate Mauriciosaurus from all other polycotylids, warranting the naming of a new genus. These include the sophisticated pattern of ridges on the bottom of the parasphenoid bone on its palate; the narrow openings in the palate bordered by the pterygoid bones; the lack of perforations in the surface of the coracoid; and the highly unusual arrangement of gastralia, or belly ribs, which is only otherwise seen in the non-polycotylid Cryptoclidus.
The Kristianstad Basin is a Cretaceous-age structural basin and geological formation in northeastern Skåne, the southernmost province of Sweden. The basin extends from Hanöbukten, a bay in the Baltic Sea, in the east to the town of Hässleholm in the west and ends with the two horsts Linderödsåsen and Nävlingeåsen in the south. The basin's northern boundary is more diffuse and there are several outlying portions of Cretaceous-age sediments. During the Cretaceous, the region was a shallow subtropical to temperate inland sea and archipelago.
Archaeolamna is an extinct genus of mackerel sharks that lived during the Cretaceous. It contains three valid species which have been found in Europe, North America, and Australia. While it is mostly known from isolated teeth, an associated set of teeth, jaws, cranial fragments, and vertebrae of A. kopingensis is known from the Pierre Shale of Kansas. Teeth of A. k. judithensis were found with a plesiosaur skeleton with bite marks from the Judith River Formation of Montana. It was a medium-sized shark with an estimated total body length of 3–4 metres (9.8–13.1 ft).
Pseudoscapanorhynchidae is a family of extinct mackerel sharks that lived during the Cretaceous and potentially the Paleogene. It currently includes Cretodus, Eoptolamna, Leptostyrax, Protolamna, Pseudoscapanorhynchus, and possibly Lilamna.