| Cyperus diandrus | |
|---|---|
| | |
| North shore of Lake Ontario, Canada in August | |
| Scientific classification | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Angiosperms |
| Clade: | Monocots |
| Clade: | Commelinids |
| Order: | Poales |
| Family: | Cyperaceae |
| Genus: | Cyperus |
| Species: | C. diandrus |
| Binomial name | |
| Cyperus diandrus | |
| Synonyms [3] | |
Homotypic synonyms
Heterotypic synonyms
| |
Cyperus diandrus, also known as umbrella flatsedge, is a species of flowering plant in the sedge family Cyperaceae. It is native to northeastern North America. It is sometimes confused with Cyperus bipartitus , both of which have chestnut-brown scales protecting the flowers. At one time, botanists considered the latter to be a variety of Cyperus diandrus.
Cyperus diandrus is an annual sedge with fibrous roots. There are multiple stems at the base of the plant. A stem rarely exceeds 25 cm (9.8 in) in height. Above the stem, the inflorescence bears clusters of spikelets, each with numerous florets. There are 2–3 leaf-like bracts at the base of the inflorescence. The main cluster of spikelets at the top of the stem is stalkless, accompanied by 0–4 additional clusters on stalks up to 6 cm long. Each cluster supports 6–12 flattened spikelets. The terminal spikelet is usually erect. A spikelet consists of 8–28 florets, each covered by a scale (a type of bract in sedges and grasses). Each floret is bisexual with 2 (occasionally 3) anthers and a single style with two branched stigmas. The anthers and stigmas emerge from the axil of the floral scale but a perianth is absent. The fruit is an achene. [4] [5]
Cyperus diandrus is sometimes confused with Cyperus bipartitus , both of which have pigmented floral scales ranging in color from reddish-brown to purplish-brown. The two species may be distinguished by the pigmented pattern on the surface of the scales. [5] [6] [7] Occasionally the scales will be entirely pale greenish-white. [8] Cyperus diandrus has conspicuous, persistent styles while those of Cyperus bipartitus are shorter and less conspicuous. The long styles of Cyperus diandrus give its spikelets a woolly appearance.
Cyperus diandrus was named and described by the American botanist John Torrey in 1819. [2] Its type specimen was collected in salt marshes near New York City. [9] Torrey referred to the species as the "diandrous galingale". [10] As of February 2026 [update] , the botanical name Cyperus diandrusTorr. is widely accepted. [3] [11] [12] [13]
In 1813, the German-American botanist Frederick Traugott Pursh described a variety of Cyperus flavescens with chestnut-brown spikelets. [14] Pursh's description was based on specimens from New York, Vermont, and Massachusetts. [15] In 1836, Torrey published the name Cyperus diandrus var. castaneus based on Pursh's variety. [16] Torrey described a taxon with cespitose habit, shining dark-chestnut scales, and relatively short styles. [17] [18] As of February 2026 [update] , the name Cyperus diandrus var. castaneus(Pursh) Torr. is considered to be a synonym of Cyperus bipartitusTorr. [19]
| Cyperus diandrus | Cyperus bipartitus (syn. Cyperus diandrus var. castaneus) | |
|---|---|---|
| Spikelets | Flatly compressed | Less so |
| Scales, shape | Oblong, with obtuse tips | Longer and narrower, very closely imbricated (overlapping) |
| Scales, pigment | Thin band of pigment along the margin, with a conspicuous pale area on the face of the scale | More heavily pigmented such that a pale area is lacking |
| Scales, texture | Membranaceous (thin, pliable) | Coriaceous (leathery), with a firmer texture |
| Styles | Much exserted beyond the scales (2–4 mm); three or four times the length of the achene; 2-cleft nearly to the base; straight | Scarcely exserted beyond the scales(<2 mm); about twice the length of the achene; less deeply cleft; usually making a short double curvature toward the base |
| Achenes | Oblong-obovate, slightly scabrous, dull | Orbicular-obovate, minutely wrinkled transversely |
Cyperus diandrus is native to northeastern North America, ranging from Ontario and Nebraska to Nova Scotia and Virginia. [20] It was introduced in the state of Washington. [21] It grows on shorelines in sandy, peaty and slightly brackish habitats, usually not in disturbed areas. [4]
Cyperus diandrus is a flowering plant whose season runs from July to October. [22]
As of February 2026 [update] , the NatureServe global conservation status of Cyperus diandrus is secure (G5). [1] It is imperiled (S2) in Pennsylvania. [23]