| Deuterosaurus Temporal range: Capitanian | |
|---|---|
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| Deuterosaurus biarmicus | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Clade: | Synapsida |
| Clade: | Therapsida |
| Suborder: | † Dinocephalia |
| Family: | † Deuterosauridae |
| Genus: | † Deuterosaurus Eichwald, 1846 |
| Type species | |
| Deuterosaurus biarmicus Eichwald, 1846 | |
| Species | |
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Deuterosaurus is an extinct genus of dinocephalian therapsids, one of the non-mammalian synapsids dominating the land during the late Paleozoic.
Deuterosaurus comes from Greek δευτερος "second" and σαυρος "lizard". [1]
Skulls of Deuterosaurus are well known from several finds. It is over 10 inches (25 cm) high, [10] with a long snout and conical teeth. Like all anteosaurs, the skull possessed long, dagger-like canine teeth. The skull was rather short for an anteosaur, with a broad cheek region, indicating a very strong bite. The eyes were partly slanted forward, giving it at least partial stereo vision. The pineal eye, though small, had a well formed opening right atop the brain case.
Deuterosaurus was a very large animal, the size of a modern grizzly bear. T. H. Huxley mistakenly considered it to be a dinosaur. [11] Judging from related therapsids, the short but massive legs were held splayed, much like a modern crocodile. When walking, the tail would have swung sideways, like in modern reptiles.
Deuterosaurus is found in what is now Siberia, which in the Permian was dominated by temperate lowlands. Deuterosaurus was among the largest animals of its day, and has variously been interpreted as a herbivore or carnivore. While the large canines may indicate the ability to kill prey, the short legs and massive body would have made it unsuited as a long distance runner, and better suited to eating plants. Then again, the possible stereoscopic vision again indicates an ambush style carnivore, and the rather blunt, cone-like post canine teeth can be interpreted both ways. Possibly Deuterosaurus was omnivorous, like a modern bear.
Deuterosaurus, like all its therapsid cousins, probably laid eggs. A remarkable thickening of the skull above the eyes indicates it may have engaged in head-butting, possibly in connection with mating or territorial disputes.
Dinosaurus is an extinct genus of therapsid of controversial affinities. Its type and only species is Dinosaurus murchisonii. It is only known from a partial snout from the Permian of Russia. Its taxonomic history is intertwined with several other poorly-known Russian therapsids, particularly Rhopalodon, Brithopus, and Phthinosuchus.
Ulemosaurus is an extinct genus of dinocephalian therapsids that lived 265 to 260 million years ago, at Isheevo in Russian Tatarstan. It was a tapinocephalid, a group of bulky herbivores which flourished in the Middle Permian. Ulemosaurus and other tapinocephalians disappeared at the end of the Middle Permian.
Anteosaurs are a group of large, primitive carnivorous dinocephalian therapsids with large canines and incisors and short limbs, that are known from the Middle Permian of South Africa, Russia, China, and Brazil. Some grew very large, with skulls 50–80 centimetres (20–31 in) long, and were the largest predators of their time. They died out at the end of the Middle Permian, possibly as a result of the extinction of the herbivorous Tapinocephalia on which they may have fed.
Anteosauridae is an extinct family of large carnivorous dinocephalian therapsids that are known from the Middle Permian of Asia, Africa, and South America.These animals were by far the largest predators of the Permian period, with skulls reaching 80 cm in length in adult individuals, far larger than the biggest gorgonopsian.
Estemmenosuchidae is an extinct family of large, very early herbivorous therapsids that flourished during the Guadalupian period. They are distinguished by horn-like structures, probably for display or agonistic behavior. Apart from the best known genus, Estemmenosuchus, the group is poorly known. To date, their fossils are known only from the Perm region of Russia.
Anteosaurus is an extinct genus of large carnivorous dinocephalian synapsid. It lived at the end of the Guadalupian during the Capitanian stage, about 265 to 260 million years ago in what is now South Africa. It is mainly known by cranial remains and few postcranial bones. Measuring 5–6 m (16–20 ft) long and weighing about 600 kg (1,300 lb), Anteosaurus was the largest known carnivorous non-mammalian synapsid and the largest terrestrial predator of the Permian period. Occupying the top of the food chain in the Middle Permian, its skull, jaws and teeth show adaptations to capture large prey like the giants titanosuchids and tapinocephalids dinocephalians and large pareiasaurs.
Ivantosaurus is an extinct genus of therapsid that lived in Russia during the Wordian stage of the Permian period, named in honor of paleontologist Ivan Antonovich Efremov.
Styracocephalus platyrhynchus is an extinct genus of dinocephalian therapsid that existed during the mid-Permian throughout South Africa, but mainly in the Karoo Basin. It is often referred to by its single known species Styracocephalus platyrhynchus. The Dinocephalia clade consisted of the largest land vertebrates and herbivores during the early to mid-Permian. This period is often also referred to as the Guadalupian epoch, approximately 270 to 260 million years ago.
Venyukovia is an extinct genus of venyukovioid therapsid, a basal anomodont from the Middle Permian of Russia. The type and sole species, V. prima, is known only by a partial lower jaw with teeth. Venyukovia has often been incorrectly spelt as 'Venjukovia' in English literature. This stems from a spelling error made by Russian palaeontologist Ivan Efremov in 1940, who mistakenly replaced the 'y' with a 'j', which subsequently permeated through therapsid literature before the mistake was caught and corrected. Venyukovia is the namesake for the Venyukovioidea, a group of small Russian basal anomodonts also including the closely related Otsheria, Suminia, Parasuminia and Ulemica, although it itself is also one of the poorest known. Like other venyukovioids, it had large projecting incisor-like teeth at the front and lacked canines, although the remaining teeth are simple compared to some other venyukovioids, but may resemble those of Otsheria.
Megawhaitsia is an extinct genus of large therocephalian therapsids who lived during the Late Permian (Wuchiapingian) in what is now Eastern Europe. The only known species is M. patrichae, described in 2008 from several fossils discovered in various oblasts of European Russia. The fossils are representative of a large animal whose skull size is estimated to be 40–50 cm (16–20 in) long.
Eurosaurus is an extinct genus of non-mammalian synapsids. Its taxonomic history is intertwined with that of the therapsids Brithopus and Orthopus and the temnospondyl Melosaurus.
Mnemeiosaurus jubilaei or Deuterosaurus jubilaei is an extinct species of dinocephalian therapsids. It was originally named as representing a distinct genus, Mnemeiosaurus, by Franz Nopcsa in 1928, which some authors, such as Boonstra and Tchudinov, have agreed with, whereas some authors, such as Efremov and Ivakhnenko, regard it as a species of Deuterosaurus. Two skulls, the holotype PIN 1954/2 and a referred specimen, PIN 2629/1, are known.
Niaftasuchus is an extinct genus of therapsids. Its type and only named species is Niaftasuchus zekkeli.
Reiszia is an extinct genus of therapsids from European Russia belonging to the family Nikkasauridae.
Akidnognathidae is an extinct family of therocephalian therapsids from the Late Permian and Early Triassic of South Africa, Russia and China. The family includes many large-bodied therocephalians that were probably carnivorous, including Moschorhinus and Olivierosuchus. One akidnognathid, Euchambersia, may even have been venomous. Akidnognathids have robust skulls with a pair of large caniniform teeth in their upper jaws. The family is morphologically intermediate between the more basal therocephalian group Scylacosauridae and the more derived group Baurioidea.
Venyukovioidea is an infraorder of anomodont therapsids related to dicynodonts from the Permian of Russia. They have also been known as 'Venjukovioidea', as well as by the similar names 'Venyukoviamorpha' or 'Venjukoviamorpha' in literature. This in part owes to a misspelling by Russian palaeontologist Ivan Efremov in 1940 when he mistakenly spelt Venyukovia, the namesake of the group, with a 'j' instead of a 'y', which permeated through subsequent therapsid literature before the mistake was caught and corrected. The order Ulemicia has also been coined for a similar taxonomic concept in Russian scientific literature, which notably excludes Suminia and Parasuminia.
Pampaphoneus is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 268 to 265 million years ago during the Wordian age of the Guadalupian period in what is now Brazil. Pampaphoneus is known by an almost complete skull with the lower jaw still articulated, discovered on the lands of the Boqueirão Farm, near the city of São Gabriel, in the state of Rio Grande do Sul. A second specimen from the same locality was reported in 2019 and 2020 but has not yet been described. It is composed of a skull associated with postcranial remains. It is the first South American species of dinocephalian to have been described. The group was previously known in South America only by a few isolated teeth and a jaw fragment reported in 2000 in the same region of Brazil. Phylogenetic analysis conducted by Cisneros and colleagues reveals that Pampaphoneus is closely related to anteosaurs from European Russia, indicating a closer faunal relationship between South America and Eastern Europe than previously thought, thus promoting a Pangea B continental reconstruction.
Sinophoneus is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 272 to 270 million years ago at the beginning of the Middle Permian in what is now the Gansu Province in northern China. It is known by a skull of an adult individual, as well as by many skulls of juvenile specimens. The latter were first considered as belonging to a different animal, named Stenocybus, before being reinterpreted as immature Sinophoneus. Sinophoneus shows a combination of characters present in other anteosaurs. Its bulbous profile snout and external nostrils located in front of the canine are reminiscent of the basal anteosaur Archaeosyodon, while its massive transverse pterygoids processes with enlarged distal ends are more similar to the more derived anteosaurs Anteosaurus and Titanophoneus. First phylogenetic analyzes identified Sinophoneus as the most basal Anteosaurinae. A more recent analysis positioned it outside the Anteosaurinae and Syodontinae subclades, and recovers it as the most basal Anteosauridae.
Novocynodon is an extinct genus of therapsids from the Middle Permian of Russia. Fossils have been found in Alexandrovsky District, Orenburg Oblast. The type and only species is Novocynodon kutorgai. It was classified as a thrinaxodontid cynodont when first described in 2012, but this was disputed by Kammerer (2014), who argued that it might instead be a juvenile anomodont or dinocephalian.
Parasumina is an extinct genus of anomodont known from the late Capitanian age at the end of the middle Permian period of European Russia. The type and only species is Parasuminia ivakhnenkoi. It was closely related to Suminia, another Russian anomodont, and was named for its resemblance. Little is known about Parasuminia as the only fossils are of fragmentary pieces of the skull and jaw, but the known remains suggest that its head and jaws were deeper and more robust than those of Suminia, and with shorter, stouter teeth. However, despite these differences they appear to have been similar animals with a similarly complex method of processing vegetation.
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