A therapsid is a member of the clade Therapsida, which is a major group of eupelycosaurian synapsids that includes mammals and their ancestors and close relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, as opposed to the sprawling posture of many reptiles and salamanders.
Dinocephalians are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania.
Biarmosuchus is an extinct genus of biarmosuchian therapsids that lived around 267 mya during the Middle Permian period. Biarmosuchus was discovered in the Perm region of Russia. The first specimen was found in channel sandstone that was deposited by flood waters originating from the young Ural Mountains.
Eotitanosuchus is an extinct genus of biarmosuchian therapsids whose fossils were found in the town of Ochyor in Perm Krai, Russia. It lived about 267 million years ago. The only species is Eotitanosuchus olsoni.
Biarmosuchia is an extinct clade of non-mammalian synapsids from the Permian. Biarmosuchians are the most basal group of the therapsids. They were moderately-sized, lightly built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation.
Inostrancevia is an extinct genus of large carnivorous therapsids which lived during the Late Permian in what is now European Russia and Southern Africa. The first-known fossils of this gorgonopsian were discovered in the Northern Dvina, where two almost complete skeletons were exhumed. Subsequently, several other fossil materials were discovered in various oblasts, and these finds will lead to a confusion about the exact number of valid species in the country, before only three of them were officially recognized: I. alexandri, I. latifrons and I. uralensis. More recent research carried out in South Africa has discovered fairly well-preserved remains of the genus, being attributed to the species I. africana. An isolated left premaxilla suggests that Inostrancevia also lived in Tanzania during the earliest Lopingian age. The whole genus is named in honor of Alexander Inostrantsev, professor of Vladimir P. Amalitsky, the paleontologist who described the taxon.
Estemmenosuchidae is an extinct family of large, very early herbivorous therapsids that flourished during the Guadalupian period. They are distinguished by horn-like structures, probably for display or agonistic behavior. Apart from the best known genus, Estemmenosuchus, the group is poorly known. To date, their fossils are known only from the Perm region of Russia.
Ennatosaurus is an extinct genus of caseid synapsid that lived during the Middle Permian in northern European Russia. The genus is only represented by its type species, Ennatosaurus tecton, which was named in 1956 by Ivan Antonovich Efremov. The species is known from at least six skulls associated with their lower jaws, as well as from the postcranial bones of several juvenile individuals. Ennatosaurus has the typical caseid skull with a short snout tilted forward and very large external nares. However, it differs from other derived caseids by its postcranial skeleton with smaller proportions compared to the size of the skull. As with other advanced caseids, the teeth of Ennatosaurus were well suited for slicing and cutting vegetation. The presence of a highly developed hyoid apparatus indicates the presence of a massive and mobile tongue, which had to work in collaboration with the palatal teeth during swallowing. With a late Roadian - early Wordian age, Ennatosaurus is one of the last known caseids.
Chroniosuchus is an extinct genus of chroniosuchid from the upper Permian period. The genus was first named by Vjuschkov in 1957.
Styracocephalus platyrhynchus is an extinct genus of dinocephalian therapsid that existed during the mid-Permian throughout South Africa, but mainly in the Karoo Basin. It is often referred to by its single known species Styracocephalus platyrhynchus. The Dinocephalia clade consisted of the largest land vertebrates and herbivores during the early to mid-Permian. This period is often also referred to as the Guadalupian epoch, approximately 270 to 260 million years ago.
Venyukovia is an extinct genus of venyukovioid therapsid, a basal anomodont from the Middle Permian of Russia. The type and sole species, V. prima, is known only by a partial lower jaw with teeth. Venyukovia has often been incorrectly spelt as 'Venjukovia' in English literature. This stems from a spelling error made by Russian palaeontologist Ivan Efremov in 1940, who mistakenly replaced the 'y' with a 'j', which subsequently permeated through therapsid literature before the mistake was caught and corrected. Venyukovia is the namesake for the Venyukovioidea, a group of small Russian basal anomodonts also including the closely related Otsheria, Suminia, Parasuminia and Ulemica, although it itself is also one of the poorest known. Like other venyukovioids, it had large projecting incisor-like teeth at the front and lacked canines, although the remaining teeth are simple compared to some other venyukovioids, but may resemble those of Otsheria.
Megawhaitsia is an extinct genus of large therocephalian therapsids who lived during the Late Permian (Wuchiapingian) in what is now Eastern Europe. The only known species is M. patrichae, described in 2008 from several fossils discovered in various oblasts of European Russia. The fossils are representative of a large animal whose skull size is estimated to be 40–50 cm (16–20 in) long.
Niaftasuchus is an extinct genus of therapsids. Its type and only named species is Niaftasuchus zekkeli.
Ulemica is an extinct genus of venyukovioid therapsids, a type of anomodont related to dicynodonts. It lived during the Middle Permian period in what is now Russia, and is known from the Isheevo assemblage of the Amanakskaya Formation. The type species, U. invisa, was originally placed in the genus Venyukovia by Russian palaeontologist Ivan Efremov in 1940. It was later given its own genus Ulemica in 1996 by Mikhaïl Ivakhnenko, who also named a second species U. efremovi. Efremov had originally intended to name the fossils of U. invisa as 'Myctosuchus invisus', however, he later recognised their similarity to Venyukovia and chose to assign the Isheevo material to this genus and leaving 'Myctosuchus' a nomen nudum.
Phaanthosaurus is an extinct genus of basal procolophonid parareptile from early Triassic deposits of Nizhnii Novgorod, Russian Federation. It is known from the holotype PIN 1025/1, a mandible. It was collected from Vetluga River, Spasskoe village and referred to the Vokhmian terrestrial horizon of the Vokhma Formation. It was first named by P. K. Chudinov and B. P. Vjushkov in 1956 and the type species is Phaanthosaurus ignatjevi.
Gorochovetzia is an extinct genus of therocephalian therapsids from the Late Permian of Russia. Fossils have been found in the Gorokhovetsky District of Vladimir Oblast. Gorochovetzia is a member of the family Hofmeyriidae. Its type and only species is G. sennikovi, named in 2011. The skull is short and very robust. The canine teeth are large, while those behind them have enlarged crowns and serrated edges. The lower jaw is deep and curved upward.
Purlovia is an extinct genus of herbivorous therocephalian therapsids from the Late Permian of Russia. Together with the closely related South African genus Nanictidops, it is a member of the family Nanictidopidae. Fossils have been found from the Tonshayevsky District of Nizhny Novgorod Oblast. The type species of Purlovia, P. maxima, was named in 2011.
Scalopodon is an extinct genus of therocephalian therapsids from the Late Permian of Russia. The type species Scalopodon tenuisfrons was named in 1999 from the Kotelnichsky District of Kirov Oblast. Scalopodon is known from a single fragmentary holotype specimen including the back of the skull, the left side of the lower jaw and isolated postorbital and prefrontal bones. The skull was found in the Deltavjatia Assemblage Zone, which dates back to the early Wuchiapingian about 260 million years ago. Distinguishing features of Scalopodon include narrow frontal bones and a distinctive sagittal crest along the parietal region at the back of the skull. Scalopodon was originally classified in the family Scaloposauridae, and was the first scaloposaurid found in Russia. More recent studies of therocephalians have found scaloposaurids like Scalopodon to be juvenile forms of larger therocephalians and do not consider Scaloposauridae to be a valid group. Scalopodon and most other scaloposaurids are now classified as basal members of Baurioidea.
Karenites is an extinct genus of therocephalian therapsids from the Late Permian of Russia. The only species is Karenites ornamentatus, named in 1995. Several fossil specimens are known from the town of Kotelnich in Kirov Oblast.
Phthinosaurus is an extinct genus of therapsids from the Middle Permian of Russia. The type species Phthinosaurus borrisiaki was named by Soviet paleontologist Ivan Yefremov in 1940 on the basis of an isolated lower jaw. Because this jaw provides few distinguishing characteristics, the evolutionary relationships of Phthinosaurus are poorly known. Yefremov named the family Phthinosuchidae in 1954 to include Phthinosaurus and the newly named Phthinosuchus, which was described on the basis of a crushed partial skull. American paleontologist Everett C. Olson placed both of these therapsids in the larger infraorder Phthinosuchia in 1961. In 1974 Leonid Tatarinov named the family Phthinosauridae to include Phthinosaurus alone, retaining Phthinosuchus within Phthinosuchidae.
